Introduction

The fauna of El Salvador has been variably studied. The richness and distribution of fish (^{González-Murcia, 2019}; ^{McMahan et al., 2013}), amphibians and reptiles (^{Köhler et al., 2006}), birds (^{Dickey & Van Rossem, 1938}; ^{Ibarra-Portillo, 2013}), and mammals (^{Owen & Girón, 2012}) have been reasonably inventoried and documented; however, the knowledge on invertebrate animals is quite different. Less than 1% of the literature on Salvadoran biodiversity is about the invertebrate animal fauna (^{Gallo, 2005}), which leaves many gaps in what we know of the distribution of the species. A clear example is Coleoptera, the most speciose order of Insecta, with at least 386,000 described species in 176 families (^{Bouchard et al., 2017}). The species richness of Coleoptera in El Salvador is unknown, but recent efforts have been made to catalog the beetle fauna. For example, ^{Van Roie et al. (2019)} documented 420 species and 132 genera of Chrysomelidae, and ^{Pablo-Cea et al. (2021)} recorded 96 especies in 38 genera of Staphylinidae. Both families are important for their diverse feeding habits and behaviors in a variety of ecosystems (^{Asenjo et al., 2013}; ^{Jolivet 1988}; ^{Navarrete-Heredia, 2002}). Also, they are taxonomically diverse families and together with Scarabaeidae, Curculionidae, Carabidae, and Cerambycidae constitute the 6 megadiverse families of Coleoptera (^{Bouchard et al., 2017}).

The objective of the present work was to document the current species list of Scarabaeoidea in El Salvador and their distributions. Scarab beetles are an important group due to their broad diversity of feeding habits and the functions the member species perform in ecosystems (^{Deloya et al., 2007}). Scarab beetles are easily recognized by the asymmetrical antennal club of multiple lamellae that are united or separated (^{Marshall et al., 2018}). The superfamily encompasses 13 extant families worldwide (^{Bouchard et al., 2017}), 10 of which occur in the New World: Geotrupidae, Glaphyridae, Glaresidae, Hybosoridae, Lucanidae, Diphyllostomatidae, Ochodaeidae, Passalidae, Pleocomidae, Scarabaeidae, and Trogidae. Worldwide, there are about 2,500 genera and 35,000 recognized especies (^{Scholtz & Grebennikov, 2005}), and in the New World there are at least 7,403 species distributed among 267 genera (^{Deloya et al., 2018}).

The first valid scarabaeoid species recorded in El Salvador date from the 19^th Century. These species, documented by the German A. F. ^{Kuwert (1897}, ¹⁸⁹⁸⁾ in “Die Passaliden dichotomisch bearbeitet”, were 5 passalids: Arrox agassizi (Kaup), Chondrocephalus granulum (Kuwert), Ogyges championi (Bates), Petrejoides subrecticornis (Kuwert), and Vindex sculptilis Bates. Thirty-seven years later, ^{Hincks and Dibb (1935)} added 2 passalid species, Paxillus leachi MacLeay and Ptichopus angulatus (Percheron), without giving a specific collection locality in the country. The decade 1950-1959 was the second most important period in the history of cataloging the Scarabaeoidea of El Salvador, when 61 species were documented among 5 publications. Two works by German authors presented important contributions to the knowledge of the scarabaeoid beetles of the country: “The Passalidae (Ins. Col.) of El Salvador” by W. D. ^{Hincks (1953)} added 7 passalid species; and “Scarabaeidae Rutelinae aus El Salvador” by E. ^{Franz (1955)} recorded 13 new country records in the subfamily. Described in this work was Paranomala vicenti (Franz), 1 of 6 precinctive scarabaeoid species in El Salvador. In 1957, one of the principal contributions to the entomological knowledge in El Salvador, “Lista de Insectos Clasificados de El Salvador” by P. A. Berry and M. Salazar-Vaquero, was published. In this technical bulletin, 29 species were added for the superfamily, mostly in the family Scarabaeidae (Scarabaeinae: 4 species, Rutelinae: 8 species, Dynastinae: 8 species, and Cetoniinae: 8 species), and 1 species of Trogidae, Omorgus suberosus (Fabricius). Of equal importance was “Segunda Lista de Insectos Clasificados de El Salvador” (^{Berry, 1959b}) that complemented the first list and added 11 more species of Scarabaeidae (2 species each of Aphodiinae, Scarabaeinae, Rutelinae, and Dynastinae, along with 3 species of Melolonthinae). At the end of the decade, ^{Vaurie (1958)} cited Diplotaxis poropyge Bates in the country.

In the 1960s and 1970s, another 26 scarab beetle species were added to the Salvadoran fauna. ^{Vaurie (1960)} added 10 species of Diplotaxis Kirby. ^{Howden and Cartwright (1963)} recorded Onthophagus incensus Say and described O. batesi. The only contributions to the knowledge of Geotrupidae are from ^{Howden (1964}, ¹⁹⁷⁴⁾, who reported 7 species from the country. ^{Howden (1974)} described Haplogeotrupes guatemalensis tridentatus (Howden), a subspecies considered precinctive to the country. ^{Cartwright (1967)} added 1 species of Aphodiinae, ^{Howden (1968)} added 3 species of Cetoniinae, and 1 species of Passalidae was documented for the first time by ^{Reyes-Castillo (1973)}. ^{Ratcliffe (1978)} recorded a second species of Trogidae, Omorgus fuliginosus Robinson. ^{Andrews et al. (1979)} recorded 2 phytophagous species of Melolonthinae and Rutelinae as crop pests.

In the 1980s, 18 additional species were reported: 1 species of Passalidae (^{Schuster, 1989}), 1 of Aphodiinae (^{Gordon & Cartwright, 1980}), 11 of Scarabaeinae (^{Howden & Young, 1981}; ^{Zunino & Halffter, 1988}), 2 of Rutelinae (^{Morón, 1983}, ¹⁹⁸⁷) , 1 of Melolonthinae (^{Morón, 1988}), and 2 of Cetoniinae (^{Cave, 1983}), including 3 precinctive species: Onthophagus salvadorensis Zunino & Halffter (Scarabaeinae), Chlaenobia solanophaga (Morón) (Melolonthinae), and Chondrocephalus salvadorae (Schuster) (Passalidae). In the 1990s, 27 species were recorded in 12 publications. ^{Schuster and Reyes-Castillo (1990)}, ^{Morón and Howden (1992)}, and ^{Warner (1992)} recorded single species of Passalidae, Melolonthinae, and Dynastinae, respectively. ^{Ratcliffe and Deloya (1992)} documented 2 species of Cetoniinae, including the precinctive Hologymnetis vulcanorum Ratcliffe & Deloya; and 8 species of Phyllophaga Harris and 2 of Chlaenobia Blanchard were added to the fauna by ^{King (1994)}, ^{Mendoza (1994)}, and ^{Morón (1994)}. Ten species of Scarabaeinae were added by ^{Edmonds (1994)}, ^{Kohlmann and Solís (1997)}, ^{Fuentes (1998)}, and ^{Rivera-Cervantes and Halffter (1999)}.

The period 2000-2009 is the most important in understanding the fauna of Scarabaeoidea in El Salvador due to the addition of 97 species. ^{Schuster and Cano (2005)} reported 1 species of Passalidae. Eight publications newly recorded 22 species of Aphodiinae, principally in the genus Ataenius Harold, with 16 species (^{Dellacasa et al., 2002}, ²⁰⁰⁸; ^{Galante et al., 2003}; ^{Horgan, 2008}; ^{Stebnicka, 2001}, ²⁰⁰², ²⁰⁰³, ²⁰⁰⁵, ²⁰⁰⁶, ²⁰⁰⁷; ^{Stebnicka & Lago, 2005}; ^{Stebnicka & Skelley, 2007}). ^{Kohlmann (2000)}, ^{Delgado and Kohlmann (2001)}, ^{Horgan (2001}, ²⁰⁰², ²⁰⁰⁸⁾, ^{Kohlmann et al. (2003)}, ^{Morón (2003)}, ^{Solís and Kohlmann (2004)}, ^{Pulido-Herrera and Zunino (2007)}, ^{Fuentes (2008}, ²⁰⁰⁹⁾, and ^{Génier (2009)} reported new records for 17 species of Scarabaeinae. ^{Morón (2006)}, ^{Arce-Pérez (2008)}, and ^{Evans and Smith (2009)} documented the first country records for 7 species of Melolonthinae, including 4 species of Macrodactylus Dejean. ^{Morón and Paucar-Cabrera (2003)} and ^{Delgado et al. (2006)} added 2 species of Rutelinae. In 2006, Ratcliffe and Cave published an important contribution to the scarab beetle knowledge in El Salvador: “The Dynastine Scarab Beetles of Honduras, Nicaragua, and El Salvador (Coleoptera: Scarabaeidae: Dynastinae)”. They inventoried the fauna of Dynastinae in El Salvador, Honduras and Nicaragua, adding 46 species to the scarab beetle fauna of the country. The only species of Hybosoridae known to occur in El Salvador, Anaides laticollis Harold and Chaetodus teamscaraborum Ocampo, were documented by ^{Ocampo (2006)}.

The decade of 2010-2019 is the third most important historical period in the development of knowledge about Salvadoran Scarabaeoidea. Thirty-one more species were newly recorded, among them, 1 species of Passalidae, Chondrocephalus granulifrons (Bates) (^{Serrano-Peraza, 2017}), 1 of Ochodaeidae, Parochodaeus puncticollis (Arrow) (^{Paulsen, 2012}), and 1 of Lucanidae, Trogellus (Mayaesalus) trifinius Paulsen, that is precinctive to the country (^{Paulsen, 2013}). Three Aphodiinae were added to the Salvadoran fauna by ^{Dellacasa et al. (2012}, ²⁰¹³⁾, including the precinctive Gonaphodioides cartwrighti Dellacasa, Dellacasa, & Gordon. Seven species of Scarabaeinae, including the non-native Digitonthophagus gazella (Fabricius), were recorded by ^{Edmonds and Zidek (2010)}, ^{Solís and Kohlmann (2013)}, ^{Deloya et al. (2014)}, ^{Pablo-Cea (2014)}, and ^{Pablo-Cea et al. (2017)}. ^{Rivera-Gasperín and Morón (2017)} newly reported 1 species of Chlaenobia, and the Salvadoran K. A. ^{Serrano-Chicas (2019)} added 4 more species of Melolonthinae and 4 species de Rutelinae. ^{López-Sorto and Sermeño-Chicas (2013)} reported for the first time 3 species of Chrysina Kirby. ^{López-Sorto et al. (2014)} reported 1 species of Dynastinae, Dynastes maya Hardy. For Cetoniinae, new country records were documented for 3 species of Euphoria Burmeister (^{Orozco, 2012}) and 2 of Gymnetis MacLeay (^{Ratcliffe, 2018}). During 2020-2022, new Salvadoran records of Melolonthinae (2 species, ^{Arce-Pérez & Morón, 2020}; ^{Pablo-Cea, 2021}), Rutelinae (1 species, ^{Pablo-Cea, 2021}), and Cetoniinae (5 species, ^{Pablo-Cea, 2021}; ^{Pablo-Cea & Alfaro, 2020}) were documented. In the catalog herein we add 19 species to the fauna of Scarabaeoidea in El Salvador: Passalidae: Ameripassalus guatemalensis (Kaup); Scarabaeidae: Aphodiinae: Ataenius carinator Harold; Melolonthinae: Diplotaxis brevipilosa Moser, D. cavifrons Moser, Isonychus pictus Sharp, Chlaenobia vexata (Horn), Phyllophaga baneta Saylo, P. colimana (Moser), and P. punctuliceps (Bates); Rutelinae: Pelidnota guatemalensis Bates, P. notata Blanchard, Paranomala championi Bates, P. plurisulcata Bates, P. semicincta Bates, P. denticollis (Bates), P. eucoma (Bates), P. sticticoptera (Blanchard), and P. undulata undulata (Melsheimer); Dynastinae: Euetheola humilis (Burmeister). We also first report the presence of the genus Germarostes Paulian (Hybosoridae: Ceratocanthinae) in the country.

Materials and methods

The compilation of data on the Scarabaeoidea known to occur in El Salvador is based on published taxonomic and ecological literature and label data with specimens available from the following collections: CMNC - Canadian Museum of Nature, Ottawa, Canada; CNCI - Canadian National Collection of Insects, Ottawa, Canada; DZUP- Coleção Entomológica Padre Jesus Santiago Moure, Departamento de Zoologia, Universidade Federal do Paraná, Curitiba, Brazil; EAPZ - Universidad Zamorano, Francisco Morazán, Honduras; EMEC - Essig Museum of Entomology, University of California, Berkeley, California, USA; FMNH - Field Museum of Natural History, Chicago, Illinois, USA; IFIT-CP - Instituto de Fitosanidad, Campus Montecillo, Colegio de Postgraduados, Mexico; MUHNES-Museo de Historia Natural de El Salvador, San Salvador, El Salvador; MZSP - Museu de Zoologia, Universidade de São Paulo, São Paulo, Brazil; RDCC - Ronald D. Cave Private Collection, Port St. Lucie, Florida, USA; TAMU-Texas A&M University, College Station, Texas, USA; UNSM - University of Nebraska State Museum, Lincoln, Nebraska, USA.

All species are organized by family, subfamily, and tribe following the classification of ^{Bouchard et al. (2011)}. The information for each species is organized as follows: Scientific name, Author, year of description.

Distribution. Primarily based on ^{Schoolmeesters (2022)}. If the species was not listed in ^{Schoolmeesters (2022)}, the reference used to establish the distribution of the species is indicated. If the species does appear in ^{Schoolmeesters (2022)} but was not cited for El Salvador, the first reference citing the species from the territory is given. The countries in the Americas are ordered north to south and west to east, followed by the West Indian countries ordered in the same way. This section does not contain an exhaustive bibliography of the complete distribution of the species, so some countries where the species occur may not be listed.

Locality records. Department: specific locality record. If the species is reported for the country only, “no specific department: no specific locality” is indicated, followed by the reference(s) and the page where it is cited for the country (as in ^{Asenjo et al., 2013}). Departments are listed from west to east as presented in Table 1 in which geocoordinates, elevation, and topographic zone are given for each locality.

Years of collection. All years in which the species has been collected (if known).

Months of collection. All months in which the species has been collected (if known). Ranges include all months within the given range.

Topographic zone. All topographic regions where the species was collected (if known).

Elevational range. Minimum to maximum elevation at which the species was collected (if known). Only 1 parameter is listed if the species is only known from a single collection site in El Salvador.

Comments. Miscellaneous observations about collection methods and the authors’ observations about the species. Old names prior to nomenclatural changes or erroneous determinations are provided if they were used for the species in older literature. Precinctive species and new records for the fauna of El Salvador are indicated. Names of localities reported in the literature or on labels with specimens deposited in museums but could not be related to any known site are noted.

A topographic map of El Salvador (Fig. 1) was used for displaying the distribution of each species with specific locality data in El Salvador. Twenty-one species have no specific locality data. The maps are presented in the same order the species are listed in the catalog. In the case of specimens with a department record only, the departmental capital’s coordinates were used for representation in the map (red points in maps).

El Salvador encompasses 21,041 km² and is the smallest country in Central America. The territory is divided into 4 well-differentiated topographic zones (^{Daugherty, 1969}): Coastal Plain (0-300 m); Volcanic Chain (250-2,200 m); Interior Valley (200-700 m); and Northern Mountains (700-2,700 m) with the highest peak in El Salvador. The Coastal Plain, Volcanic Chain, and Interior Valley belong to the Pacific Lowlands, whereas the Northern Mountains are a continuation of the Chiapas Highlands biogeographic province (^{Morrone, 2014}). El Salvador is divided into 14 departments (Fig. 1).

Results

In El Salvador, 295 species in 106 genera of scarabaeoid beetles are known to occur.

Superfamily Scarabaeoidea Latreille, 1802

Family Geotrupidae Latreille, 1802

Subfamily Bolboceratinae Mulsant, 1842

Tribe Athyreini Lynch Arribálzaga, 1878

Neoathyreus Howden & Martínez, 1963

Neoathyreus (Neoathyreus) excavatus (Laporte, 1840)

Distribution: Mexico, Guatemala, El Salvador, Honduras, Costa Rica, Panama, Colombia, Venezuela, Trinidad & Tobago, Guyana, Suriname, French Guiana, Brazil, and Peru (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (RDCC); Santa Ana: Parque Nacional Montecristo (Bosque Nebuloso) (^{Howden, 1964: 19}); San Jacinto (^{Howden, 1964: 19}); San Salvador: San Salvador (^{Howden, 1964: 19}); La Paz: Comalapa (La Providencia) (^{Horgan, 2002: 30}) (Fig. 2).

Years of collection: 1925, 1958, 1960, 1979, 1995-1998.

Months of collection: May and June.

Topographic zone: northern mountains, interior valley, coastal mountains, coastal plain.

Elevation range: 40-2,360 m.

Comments: reported as Neoathyreus quadridentatus by ^{Howden (1964)} and ^{Horgan (2002)}. Specimens were captured at lights in San Salvador (^{Howden, 1964}) and in pitfall traps baited with cow dung in Comalapa (^{Horgan, 2002}).

Neoathyreus (Neoathyreus) interruptus^{Howden, 1964}Distribution: Mexico, Guatemala, and El Salvador

(^{Schoolmeesters, 2022}).

Locality records: San Salvador: San Salvador (^{Howden, 1964: 23}) (Fig. 2).

Years of collection: 1958.

Months of collection: June.

Topographic zone: interior valley.

Elevation range: 685 m.

Neoathyreus (Neoathyreus) mexicanus (Klug, 1845) Distribution: Mexico, Guatemala, El Salvador, Nicaragua, Costa Rica, and Panama (^{Schoolmeesters, 2022}).

Locality records: Santa Ana: Parque Nacional Montecristo (Los Planes) (^{Howden, 1964: 27});

Chalatenango: Las Ceibas [probably Las Vueltas] (^{Howden, 1964: 27}); San Salvador: San Salvador (^{Howden, 1964: 27}); La Paz: Comalapa (La Providencia) (^{Horgan, 2008: 2969} ) (Fig. 2).

Years of collection: 1960, 1999.

Months of collection: May-July, and October.

Topographic zone: northern mountains, interior valley, coastal plain.

Elevation range: 40-1,890 m.

Comments: specimens from Comalapa were captured in pitfall traps baited with beef carrion (^{Horgan, 2008}).

Tribe Bolboceratini Mulsant, 1842 Bolbelasmus Boucomont, 1911

Bolbelasmus (Bolbelasmus) arcuatus (Bates, 1887) Distribution: Mexico, Guatemala, El Salvador,

Honduras, Nicaragua, Costa Rica, and Panama (^{Schoolmeesters, 2022})

Locality records: Santa Ana: San Jacinto (^{Howden, 1964: 40}); La Libertad: Parque Nacional Walter Thilo Deininger (^{Horgan, 2008: 2968}); San Salvador: San Salvador (^{Howden, 1964: 40}) (Fig. 2).

Years of collection: 1925, 1958-1960.

Months of collection: May and June.

Topographic zone: interior valley, coastal plain.

Elevation range: 135-685 m.

Comments: specimens were captured at lights in San Salvador (^{Howden, 1964}) and in pitfall traps baited with cow dung in Comalapa (^{Horgan, 2008}).

Bolbelasmus (Bolbelasmus) monticolus Howden, 1974 Distribution: Guatemala and El Salvador

(^{Schoolmeesters, 2022}).

Locality records: Santa Ana: Parque Nacional Cerro Verde (^{Howden, 1974: 569}); Parque Nacional Montecristo (Bosque Nebuloso) (RDCC); La Libertad: Volcán de San Salvador (El Boquerón) (^{Howden, 1974: 568}) (Fig. 2).

Years of collection: 1965-1967, 1971, 1999.

Months of collection: May and August.

Topographic zone: northern mountains, coastal mountains.

Elevation range: 1,845-2,360 m.

Comments: captured at light in El Boquerón.

Subfamily Geotrupinae Latreille, 1802

Tribe Geotrupini Latreille, 1802

Haplogeotrupes Nikolajev, 1979

Haplogeotrupes guatemalensis tridentatus (^{Howden, 1974})

Distribution: El Salvador (^{Schoolmeesters, 2022}).

Locality records: Santa Ana: Parque Nacional Montecristo (Bosque Nebuloso) (^{Howden, 1974: 571}) (Fig. 2).

Years of collection: 1971.

Months of collection: May.

Topographic zone: northern mountains.

Elevation range: 2,360 m.

Comments: precinctive to El Salvador. Reported as Geotrupes guatemalensis tridentatus by ^{Howden (1974)}. Specimens were collected from horse dung.

Onthotrupes Howden, 1964 Onthotrupes onitidipes (Bates, 1887)

Distribution: Mexico, Guatemala, and El Salvador (^{Schoolmeesters, 2022}).

Locality records: Santa Ana: Parque Nacional Montecristo (Bosque Nebuloso) (^{Howden, 1964: 62}) (Fig. 2).

Years of collection: 1958, 1999.

Months of collection: May.

Topographic zone: northern mountains.

Elevation range: 2,360 m.

Comments: reported as Geotrupes (Onthotrupes) onitidipes by ^{Howden (1964)}. Specimens were found under cow dung.

Family Passalidae Leach, 1815

Subfamily Passalinae Leach, 1815

Tribe Proculini Kaup, 1868

Arrox Zang, 1905

Arrox agassizi (Kaup, 1871)

Distribution: Mexico, Guatemala, El Salvador, Honduras, Nicaragua, and Costa Rica (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Apaneca (^{Boucher, 2005: 391}); Laguna Las Ninfas (MUHNES); Santa Ana: Chalchuapa (^{Boucher, 2005: 391}); Parque Nacional Cerro Verde (^{Boucher, 2005: 391}); Las Cruces (^{Boucher, 2005: 391}); Parque Nacional Montecristo (Los Planes) (^{Schuster & Cano, 2005: 258} ; ^{Serrano-Peraza, 2017: 42}; ^{Serrano-Peraza et al., 2022: 4}); La Libertad: Volcán de San Salvador (El Boquerón) (^{Boucher, 2005: 391}); San Salvador: San Salvador (^{Kuwert, 1897: 303}; ^{Hincks & Dibb, 1935: 23}; ^{Boucher, 2005: 388}); Morazán: Cerro Cacahuatique (^{Hincks, 1953: 33}; ^{Boucher, 2005: 391}); Hacienda San Pedro (MUHNES); Osicala (^{Boucher, 2005: 391}) (Fig. 2).

Years of collection: 1897, 1919, 1951, 1959, 1974, 1979, 1992, 2015.

Months of collection: June, July, and September-December.

Topographic zone: northern mountains, interior valley, coastal mountains.

Elevation range: 180-2,020 m.

Comments: reported as Sertorius assmanni Kuwert by ^{Kuwert (1897)} and ^{Hincks & Dibb (1935)}.

Chondrocephalus Kuwert, 1896

Chondrocephalus gemmae Reyes-Castillo & Castillo, 1986

Distribution: Mexico and El Salvador (^{Serrano-Peraza, 2017}; ^{Schoolmeesters, 2022}).

Locality records: Santa Ana: Parque Nacional Montecristo (Bosque Nebuloso, Los Planes) (^{Schuster & Cano, 2005: 258}; ^{Serrano-Peraza, 2017: 42}; ^{Serrano-Peraza et al., 2022: 4}) (Fig. 2).

Years of collection: 2015.

Months of collection: September-December.

Topographic zone: northern mountains.

Elevation range: 1,890-2,360 m. Chondrocephalus granulum Kuwert, 1897

Distribution: Mexico, Guatemala, and El Salvador (^{Schoolmeesters, 2022}).

Locality records: Santa Ana: Parque Nacional Montecristo (Bosque Nebuloso) (^{Hincks, 1953: 33}; ^{Reyes-Castillo, 1970: 104}); San Salvador: San Salvador (^{Kuwert, 1897: 302}; ^{Hincks & Dibb, 1935: 33}) (Fig. 2).

Years of collection: 1897, 1951.

Months of collection: June and August.

Topographic zone: northern mountains, interior valley.

Elevation range: 685-2,360 m.

Comments: larvae were collected in June and adults in August (^{Hincks, 1953}).

Chondrocephalus granulifrons (Bates, 1886) Distribution: Mexico, Guatemala, El Salvador, and

Ecuador (^{Serrano-Peraza, 2017}; ^{Schoolmeesters, 2022}). Locality records: Santa Ana: Parque Nacional

Montecristo (Bosque Nebuloso) (^{Serrano-Peraza, 2017: 42}; ^{Serrano-Peraza et al., 2022: 4}) (Fig. 3).

Years of collection: 2015.

Months of collection: September-December.

Topographic zone: northern mountains.

Elevation range: 2,360 m. Chondrocephalus salvadorae (^{Schuster, 1989})

Distribution: El Salvador (^{Schoolmeesters, 2022}).

Locality records: Santa Ana: Parque Nacional Montecristo (Bosque Nebuloso) (^{Schuster, 1989: 695}; ^{Schuster & Cano, 2005: 258}; ^{Serrano-Peraza, 2017: 44}) (Fig. 3).

Years of collection: 1960, 1972, 2015.

Months of collection: April, July, and September-December.

Topographic zone: northern mountains.

Elevation range: 2,360 m.

Comments: precinctive to El Salvador. Reported as Petrejoides salvadorae by ^{Schuster (1989)} and ^{Serrano-Peraza (2017)}.

Heliscus Zang, 1905

Heliscus eclipticus (Truqui, 1857)

Distribution: Mexico, Guatemala, El Salvador, Costa Rica, and Colombia (^{Schoolmeesters, 2022}).

Locality records: Santa Ana: Parque Nacional Montecristo (San José Ingenio, Los Planes) (^{Hincks, 1953: 32}; ^{Serrano-Peraza, 2017: 44}; ^{Serrano-Peraza et al., 2022: 4}); Finca San Jorge (^{Hincks, 1953: 32}) (Fig. 3).

Years of collection: 1951, 1973-1977, 2015.

Months of collection: April, May, July, and September-December.

Topographic zone: northern mountains, interior valley.

Elevation range: 480-1,890 m.

Comments: reported as Popilius eclipticus by ^{Hincks (1953)} and ^{Serrano-Peraza (2017)}.

Odontotaenius Kuwert, 1896

Odontotaenius striatopunctatus (Percheron, 1835) Distribution: Mexico, Belize, El Salvador, Nicaragua,

Costa Rica, Honduras, and Colombia (^{Hincks, 1953}; ^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (La Fincona, San Benito) (MUHNES, RDCC); Santa Ana: Parque Nacional Montecristo (San José Ingenio) (^{Serrano-Peraza, 2017: 43}; ^{Serrano-Peraza et al., 2022: 4}); Sonsonate: Izalco (Los Guates Farm) (MUHNES); San Salvador: Los Planes de Renderos (MUHNES); Parque Saburo Hirao (MUHNES); San Salvador (^{Hincks, 1953: 32}; ^{Reyes-Castillo, 2003: 164}); Cuscatlán: no specific locality (MUHNES) (Fig. 3).

Years of collection: 1951, 1974-1979, 2013-2015.

Months of collection: May, June, September, November, and December.

Topographic zone: northern mountains, interior valley, coastal mountains, coastal plain.

Elevation range: 470-1,005 m.

Comments: reported as Popilius striatopunctatus by ^{Hincks (1953)}. Some specimens in MHUNES are labeled as collected in Cuscatlán: La Burrada, but we are not aware of a site with that name in El Salvador.

Ogyges Kaup, 1871

Ogyges championi (Bates, 1886)

Distribution: Mexico, Guatemala, El Salvador?, and Nicaragua? (Schuster & Reyes-Castillo, 1990).

Locality records: San Salvador: San Salvador (^{Kuwert, 1897: 292}; ^{Schuster & Reyes-Castillo, 1990: 32-33}) (Fig. 3).

Years of collection: 1877.

Months of collection: December.

Topographic zone: interior valley.

Elevation range: 685 m.

Comments: reported as Ogyges laevior (Kaup) by ^{Kuwert (1897)} and ^{Schuster & Reyes-Castillo (1990)}. The presence of the species in El Salvador and Nicaragua requires confirmation (^{Schuster & Reyes-Castillo, 1990}). Ogyges hondurensis^{Schuster & Reyes-Castillo, 1990}

Distribution: El Salvador and Honduras (^{Schoolmeesters, 2022}).

Locality records: Chalatenango: Cerro El Pital (^{López-Sorto et al., 2014: 42}); La Libertad: Volcán de San Salvador (El Boquerón) (^{Schuster & Reyes-Castillo, 1990: 17}; ^{Schuster et al., 2005: 130}) (Fig. 3).

Years of collection: 1979.

Months of collection: February.

Topographic zone: northern mountains, coastal mountains.

Elevation range: 1,845-2,660 m.

Ogyges politus (^{Hincks, 1953})

Distribution: Guatemala and El Salvador (^{Schoolmeesters, 2022}).

Locality records: Santa Ana: Parque Nacional Montecristo (Bosque Nebuloso) (^{Hincks, 1953: 32}; ^{Schuster & Reyes-Castillo, 1990: 22}; ^{Schuster & Cano, 2005: 258}; ^{Cano, 2017: 9}; ^{Serrano-Peraza, 2017: 43}; ^{Cano et al., 2018: 109}; ^{Serrano-Peraza et al., 2022: 4}); Chalatenango: no specific locality [probably Cerro El Pital] (^{Cano et al., 2018: 109}) (Fig. 3).

Years of collection: 1951, 1960, 1970-1977, 1999, 2015.

Months of collection: February, March, May, July, and September-December.

Topographic zone: northern mountains.

Elevation range: 2,360 m.

Comments: reported as Procolejus politus by ^{Hincks (1953)}.

Oileus Kaup, 1869

Oileus sargi (Kaup, 1871)

Distribution: Mexico, Guatemala, El Salvador, and Costa Rica (^{Schoolmeesters, 2022}).

Locality records: Santa Ana: Parque Nacional Montecristo (Bosque Nebuloso) (^{Schuster & Cano, 2005: 258}) (Los Planes) (^{Hincks, 1953: 32}; ^{Serrano-Peraza et al., 2022: 4}) (Fig. 3).

Years of collection: 1951, 1973, 2015.

Months of collection: May, June, and September-December.

Topographic zone: northern mountains.

Elevation range: 1,890-2,360 m.

Petrejoides Kuwert, 1896

Petrejoides subrecticornis (^{Kuwert, 1897})

Distribution: Guatemala, El Salvador, and Costa Rica (^{Castillo & Reyes-Castillo, 1984})

Locality records: San Salvador: San Salvador (^{Kuwert, 1897: 301}) (Fig. 3).

Years of collection: 1877.

Months of collection: December.

Topographic zone: interior valley.

Elevation range: 685 m.

Comments: reported as Popilius scutellopunctatus Kuwert by ^{Kuwert (1897)}.

Verres Kaup, 1871

Verres hageni Kaup, 1871

Distribution: Mexico, Guatemala, El Salvador, Honduras, Nicaragua, Costa Rica, and Colombia (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Concepción de Ataco (MUHNES); Parque Nacional El Imposible (Cerro Campana) (MUHNES); (San Benito) (RDCC); Santa Ana: Finca San Jorge (^{Hincks, 1953: 33}); Parque Nacional Montecristo (San José Ingenio) (^{Serrano-Peraza, 2017: 44}; ^{Serrano-Peraza et al., 2022: 4}); Morazán: Cerro Cacahuatique (MUHNES) (Fig. 3).

Years of collection: 1951, 1975, 1979, 1999.

Months of collection: April-June.

Topographic zone: northern mountains, interior valley, coastal mountains.

Elevation range: 480-1,400 m.

Vindex Kaup, 1871

Vindex sculptilis Bates, 1886

Distribution: Mexico, Guatemala, and El Salvador (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Francisco Menéndez) (MUHNES); Santa Ana: Parque Nacional Montecristo (Bosque Nebuloso) (^{Hincks, 1953: 34}; ^{Schuster & Cano, 2005: 258}; ^{Serrano-Peraza, 2017: 44}; ^{Serrano-Peraza et al., 2022: 4}) (Los Planes) (^{Schuster & Cano, 2005: 258}); San Salvador: San Salvador (^{Kuwert, 1898: 15}; ^{Hincks & Dibb, 1935: 33}) (Fig. 4).

Years of collection: 1897, 1951, 1976.

Months of collection: June and August.

Topographic zone: northern mountains, interior valley, coastal plain.

Elevation range: 250-2,360 m.

Comments: reported as Vindex quadrangulifrons Kuwert by ^{Kuwert (1898)} and ^{Hincks & Dibb (1935)}.

Tribe Passalini Leach, 1815

Ameripassalus Jiménez-Ferbans & Reyes-Castillo, 2014 Ameripassalus guatemalensis (Kaup, 1869)

Distribution: Mexico, Guatemala, Honduras, Nicaragua (^{Schoolmeesters, 2022}), and El Salvador.

Locality records: Santa Ana: Parque Nacional Montecristo (San José Ingenio) (MUHNES) (Fig. 4).

Years of collection: no data.

Months of collection: no data.

Topographic zone: northern mountains.

Elevation range: 810 m.

Comments: new country record.

Passalus Fabricius, 1792

Passalus interstitialis Eschscholtz, 1829

Distribution: Mexico, El Salvador, Honduras, Nicaragua, Costa Rica, Colombia, Venezuela, Trinidad & Tobago, Guyana, Suriname, French Guiana, Ecuador, Brazil, Peru, Bolivia, Paraguay, Argentina, Cuba, Jamaica, and Grenada (^{Reyes-Castillo 1973}, ^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (RDCC); (San Francisco Menéndez) (MUHNES); Santa Ana: Lago de Coatepeque (Isla Teopán) (MUHNES); El Congo (RDCC); La Libertad: El Barillo (MUHNES); Los Chorros (MUHNES, RDCC); Puerto de La Libertad (^{Romero-Nápoles, 2021}); no specific department: no specific locality (^{Reyes-Castillo, 1973: 1582}) (Fig. 4).

Years of collection: 1973-1980, 1999.

Months of collection: February, March, May, and September.

Topographic zone: coastal mountains, coastal plain.

Elevation range: 30-820 m.

Passalus punctatostriatus (Percheron, 1835) Distribution: USA, Mexico, Guatemala, El Salvador, Honduras, Nicaragua, Costa Rica, Panama, Colombia, Venezuela, Suriname, and Brazil (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Laguna Las Ninfas (MUHNES); Santa Ana: Finca San Jorge (^{Hincks, 1953: 34}); Parque Nacional Montecristo (San José Ingenio) (^{Serrano-Peraza, 2017: 43}; ^{Serrano-Peraza et al., 2022: 4}); Sonsonate: Izalco (Los Guates Farm) (MUHNES); La Libertad: Parque Nacional Parque Nacional Walter Thilo Deininger (MUHNES); San Salvador: San Salvador (^{Hincks, 1953: 34}); San Vicente: Volcán de San Vicente-Chinchontepec (Hacienda El Carmen) (^{Hincks, 1953: 34}) (Fig. 4).

Years of collection: 1951, 1973-1976.

Months of collection: April-June, and November.

Topographic zone: northern mountains, interior valley, coastal mountains, coastal plain.

Elevation range: 135-2,000 m.

Passalus punctiger Le Peletier & Audinet-Serville, 1825

Distribution: USA, Mexico, Guatemala, El Salvador, Nicaragua, Costa Rica, Colombia, Venezuela, Trinidad & Tobago, French Guiana, Ecuador (Galápagos Islands), Brazil, Peru, Bolivia, Chile, Paraguay, Argentina, Jamaica, St. Vincent & the Grenadines, and Grenada (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (MUHNES, RDCC); Santa Ana: Finca San Jorge (^{Hincks, 1953: 34}); Lago de Coatepeque (Isla Teopán) (MUHNES); Parque Nacional Montecristo (San José Ingenio) (^{Serrano-Peraza, 2017: 43}; ^{Serrano-Peraza et al., 2022: 4}); La Libertad: (Laguna de) Zapotitán (^{Hincks, 1953: 34}); Parque Nacional Walter Thilo Deininger (MUHNES); San Salvador: Lago de Ilopango (Isla Asino) (MUHNES); Los Planes de Renderos (MUHNES); Parque Saburo Hirao (MUHNES); San Salvador (^{Hincks, 1953: 35}) (Fig. 4).

Years of collection: 1951, 1973-1979, 1999.

Months of collection: February, April-June, and October-December.

Topographic zone: northern mountains, interior valley, coastal mountains, coastal plain.

Elevation range: 135-1,005 m.

Comments: the specimens collected in Zapotitán and Finca San Jorge were identified as Passalus interruptus (Linnaeus) in ^{Hincks (1953)} (see species that do not occur in El Salvador).

Paxillus MacLeay, 1819 Paxillus leachi MacLeay, 1819

Distribution: Canada, USA, Mexico, Guatemala, Belize, El Salvador, Honduras, Nicaragua, Costa Rica, Panama, Colombia, Venezuela, Guyana, Suriname, French Guiana, Ecuador, Brazil, Peru, Bolivia, Chile, Paraguay, Argentina, and Uruguay (^{Schoolmeesters, 2022}). The species is not present in the Antilles (^{Jiménez-Ferbans et al., 2015}).

Locality records: Santa Ana: El Congo (MUHNES, RDCC); Finca San Jorge (^{Hincks, 1953: 34}); La Libertad: Los Chorros (RDCC); Parque Nacional Walter Thilo Deininger (MUHNES); Plan de La Laguna (MUHNES); Puerto de La Libertad (^{Romero-Nápoles, 2021}); no specific department: no specific locality (^{Hincks & Dibb, 1935: 36}) (Fig. 4).

Years of collection: 1951, 1966, 1974-1980.

Months of collection: January, February, April, May, July, and September-December.

Topographic zone: northern mountains, interior valley, coastal mountains, coastal plain.

Elevation range: 30-820 m.

Ptichopus Kaup, 1869

Ptichopus angulatus (Percheron, 1835)

Distribution: Mexico, Guatemala, El Salvador, Honduras, Nicaragua, Costa Rica, and Panama (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (MUHNES, RDCC); Santa Ana: Finca San Jorge (MUHNES); Parque Nacional Montecristo (San José Ingenio) (MUHNES); San Salvador: San Salvador (^{Hincks, 1953: 35}); no specific department: no specific locality (^{Hincks & Dibb, 1935: 63}) (Fig. 4).

Years of collection: 1951, 1975-1976, 1999.

Months of collection: April-June.

Topographic zone: northern mountains, interior valley, coastal mountains.

Elevation range: 480-810 m.

Family Trogidae MacLeay, 1819 Subfamily Omorginae Nikolajev, 2005 Omorgus Erichson, 1847

Omorgus (Omorgus) fuliginosus Robinson, 1941 Distribution: USA, Mexico, El Salvador, and Costa

Rica (^{Schoolmeesters, 2022}).

Locality records: La Libertad: La Libertad (^{Ratcliffe, 1978: 301}); Santa Tecla (^{Ratcliffe, 1978: 301}) (Fig. 4).

Years of collection: 1971.

Months of collection: May

Topographic zone: coastal mountains, coastal plain.

Elevation range: 70-900 m.

Comments: reported as Trox fuliginosus by ^{Ratcliffe (1978)}.

Omorgus (Omorgus) suberosus (Fabricius, 1775) Distribution: USA, Mexico, El Salvador, Panama,

Colombia, Venezuela, Guyana, French Guiana, Ecuador, Brazil, Peru, Bolivia, Paraguay, Argentina, Uruguay, and Martinique; and Palearctic, Oriental, and Australian regions (^{Berry & Salazar-Vaquero, 1957}; ^{Schoolmeesters, 2022}).

Locality records: Usulután: Alegría (^{Berry & Salazar-Vaquero, 1957: 47}) (Fig. 4).

Years of collection: no data.

Months of collection: no data.

Topographic zone: coastal mountains.

Elevation range: 1,130 m.

Comments: reported as Trox suberosus by ^{Berry and Salazar-Vaquero (1957)}.

Family Lucanidae Latreille, 1804 Subfamily Aesalinae MacLeay, 1819 Tribe Aesalini MacLeay, 1819 Trogellus Paulsen, 2013 Trogellus (Mayaesalus) trifinius Paulsen, 2013 Distribution: El Salvador (^{Paulsen, 2013}).

Locality records: Santa Ana: Parque Nacional Montecristo (Bosque Nebuloso) (^{Paulsen, 2013: 17}) (Fig. 4).

Years of collection: 2002.

Months of collection: June.

Topographic zone: northern mountains.

Elevation range: 2,360 m.

Comments: the species is precinctive to El Salvador.

Family Ochodaeidae Mulsant & Rey, 1871 Subfamily Ochodaeinae Mulsant & Rey, 1871 Tribe Ochodaeini Mulsant & Rey, 1871 Parochodaeus Nikolajev, 1995 Parochodaeus puncticollis (Arrow, 1911)

Distribution: Guatemala. El Salvador, and Honduras (^{Paulsen, 2012}).

Locality records: Santa Ana: no specific locality (^{Paulsen, 2012: 178}) (Fig. 5).

Years of collection: no data.

Months of collection: no data.

Topographic zone: no data.

Elevation range: no data.

Comments: the species is known only from montane forests from Chiapas to Honduras (^{Paulsen, 2012}), therefore the locality record is probably Santa Ana: Parque Nacional Montecristo (Bosque Nebuloso) as for the lucanid T. trifinius (^{Paulsen, 2013}).

Family Hybosoridae Erichson, 1847 Subfamily Anaidinae Nikolajev, 1996 Anaides Westwood, 1845 Anaides laticollis Harold, 1863

Distribution: Mexico, Guatemala, Belize, El Salvador, Honduras, Nicaragua, Costa Rica, and Panama (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (J. Pablo, personal observation); no specific department: no specific locality (^{Ocampo, 2006: 43}) (Fig. 5).

Years of collection: 2013.

Months of collection: June-November.

Topographic zone: coastal mountains.

Elevation range: 780 m.

Comments: specimens from Parque Nacional El Imposible were captured in pitfall traps baited with human dung and chicken carrion (J. Pablo, personal observation).

Chaetodus Westwood, 1845

Chaetodus (Chaetodus) teamscaraborum Ocampo, 2006 Distribution: Mexico, Guatemala, Belize, El Salvador, Honduras, Nicaragua, Costa Rica, and Panama (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (^{Ocampo, 2006: 140}) (Fig. 5).

Years of collection: no data.

Months of collection: no data.

Topographic zone: coastal mountains.

Elevation range: 780 m.

Subfamily Ceratocanthinae Martínez, 1968 Tribe Ceratocanthini Martínez, 1968 Germarostes Paulian, 1982

Comments: specimens of this genus were collected (2018-2019) in El Salvador at lights in Ahuachapán: Parque Nacional El Imposible (San Benito); Santa Ana: Parque Nacional Montecristo (San José Ingenio); La Libertad: Parque El Bicentenario (Fig. 5). Unfortunately, none of them have been identified to species level. They are deposited in the CMNC with the hope that their identity will be revealed in the future.

Family Scarabaeidae Latreille, 1802 Subfamily Aphodiinae Leach, 1815 Tribe Aphodiini Leach, 1815

Agrilinellus Dellacasa, ^{Dellacasa, & Gordon, 2008} Agrilinellus chiapasensis (^{Galante, Stebnicka & Verdú, 2003})

Distribution: Mexico and El Salvador (^{Dellacasa et al., 2008}; ^{Schoolmeesters, 2022}).

Locality records: Santa Ana: Parque Nacional Montecristo (Bosque Nebuloso) (^{Dellacasa et al., 2008: 10}) (Fig. 5).

Years of collection: 1958, 1971.

Months of collection: May.

Topographic zone: northern mountains.

Elevation range: 2,360 m.

Blackburneus Schmidt, 1913 Blackburneus charmionus (Bates, 1887)

Distribution: Mexico, El Salvador, Costa Rica, Panama, and Brazil (^{Schoolmeesters, 2022}).

Locality records: Santa Ana: Parque Nacional Cerro Verde (^{Dellacasa et al., 2002: 202}) (Fig. 5).

Years of collection: 1958.

Months of collection: June.

Topographic zone: coastal mountains.

Elevation range: 2,020 m. Blackburneus diminutus (Bates, 1887)

Distribution: Mexico, Guatemala, and El Salvador (^{Schoolmeesters, 2022}).

Locality records: Santa Ana: Parque Nacional Cerro Verde (^{Dellacasa et al., 2002: 205}); San Miguel: Laguna Olomega (^{Dellacasa et al., 2002: 205}) (Fig. 5).

Years of collection: 1958.

Months of collection: June.

Topographic zone: coastal mountains, coastal plain.

Elevation range: 80-2,020 m. Blackburneus guatemalensis (Bates, 1887) Distribution: Mexico, Guatemala, and El Salvador (^{Schoolmeesters, 2022}).

Locality records: Santa Ana: Parque Nacional Cerro Verde (^{Dellacasa et al., 2002: 195}) (Fig. 5).

Years of collection: 1958.

Months of collection: June.

Topographic zone: coastal mountains.

Elevation range: 2,020 m.

Gonaphodiellus Schmidt, 1913 Gonaphodiellus bimaculosus (Schmidt, 1909)

Distribution: Mexico and El Salvador (^{Schoolmeesters, 2022}).

Locality records: Santa Ana: Parque Nacional Montecristo (Bosque Nebuloso) (^{Dellacasa et al., 2012: 6}) (Fig. 5).

Years of collection: 1971.

Months of collection: May.

Topographic zone: northern mountains.

Elevation range: 2,360 m. Gonaphodiellus sexguttatus (Schmidt, 1916)

Distribution: Mexico, El Salvador, Honduras, Costa Rica, Panama, and Colombia (^{Schoolmeesters, 2022}).

Locality records: Santa Ana: Parque Nacional Montecristo (Bosque Nebuloso) (^{Dellacasa et al., 2012: 14}) (Fig. 5).

Years of collection: 1958, 1971.

Months of collection: May.

Topographic zone: northern mountains.

Elevation range: 2,360 m.

Gonaphodioides Dellacasa, Dellacasa, & Gordon, 2012 Gonaphodioides cartwrighti Dellacasa, Dellacasa & Gordon, 2013

Distribution: El Salvador (^{Schoolmeesters, 2022}).

Locality records: Santa Ana: Parque Nacional Montecristo (Bosque Nebuloso) (^{Dellacasa et al., 2013: 1-2}) (Fig. 6).

Years of collection: 1958.

Months of collection: May.

Topographic zone: northern mountains.

Elevation range: 2,360 m.

Comments: the species is precinctive to El Salvador.

Haroldiellus Gordon & Skelley, 2007 Haroldiellus sallei (Harold, 1863)

Distribution: USA, Mexico, Guatemala, El Salvador, Nicaragua, and Costa Rica (^{Berry, 1959b}; ^{Dellacasa et al., 2002}; ^{Schoolmeester 2022}).

Locality records: Santa Ana: no specific locality (^{Gordon & Skelley, 2022: 3}); Chalatenango: Cerro El Pital (EAPZ); La Libertad: no specific locality (^{Skelley & Keller, 2022: 3}); San Salvador: no specific locality (^{Skelley & Keller, 2022: 3}); Cuscatlán: no specific locality (^{Skelley & Keller 2022: 3}); San Vicente: no specific locality (^{Skelley & Keller, 2022: 3}) (Fig. 6); no specific department: no specific locality (^{Berry, 1959b: 7}).

Years of collection: 1994.

Months of collection: July.

Topographic zone: interior valley, Northern mountains.

Elevation range: 2,660 m.

Comments: reported as Aphodius sallaei in ^{Berry (1959b)}.

Nialaphodius Kolbe, 1908 Nialaphodius nigrita (Fabricius, 1801)

Distribution: USA, Mexico, Belize, El Salvador, Honduras, Panama, Colombia, Venezuela, Ecuador, Brazil, Peru, Chile, Paraguay, Puerto Rico, Martinique, and St. Vincent & the Grenadines; and Afrotropical region (^{Horgan, 2008}; ^{Schoolmeesters, 2022}).

Locality records: La Libertad: Parque Nacional Walter Thilo Deininger (^{Horgan, 2008: 2969}); La Paz: Comalapa (La Providencia) (^{Horgan, 2008: 2969}) (Fig. 6).

Years of collection: 1995-1997.

Months of collection: no data.

Topographic zone: coastal plain.

Elevation range: 40-135 m.

Comments: specimens were captured in baited pitfall traps in Comalapa (cow dung and fruit) and Parque Nacional Walter Thilo Deininger (cow dung) (^{Horgan, 2008}).

Tribe Eupariini Schmidt, 1910 Ataenius Harold, 1867 Ataenius aequalis Harold, 1880

Distribution: USA, Mexico, Guatemala, Belize, El Salvador, Honduras, Costa Rica, Panama, Colombia, Venezuela, Trinidad & Tobago, Guyana, Brazil, Bolivia, Argentina, Guadeloupe, and St. Vincent & the Grenadines (^{Schoolmeesters, 2022}).

Locality records: Santa Ana: Volcán San Diego (^{Stebnicka, 2005: 105}) (Fig. 6).

Years of collection: no data.

Months of collection: no data.

Topographic zone: interior valley.

Elevation range: 760. Ataenius carinator Harold, 1874

Distribution: USA, Mexico, Honduras, Costa Rica, Panama, Colombia, Venezuela, Guyana, Suriname, Ecuador, and Peru (^{Stebnicka, 2007}; ^{Schoolmeesters, 2022}), and El Salvador.

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (RDCC) (Fig. 6).

Years of collection: 2000.

Months of collection: May.

Topographic zone: coastal mountains.

Elevation range: 780 m.

Comments: new country record.

Ataenius castaniellus Bates, 1887

Distribution: Mexico, Guatemala, El Salvador, and Honduras (^{Schoolmeesters, 2022}).

Locality records: La Libertad: Los Chorros (^{Stebnicka, 2005: 110}) (Fig. 6).

Years of collection: 1971.

Months of collection: May.

Topographic zone: coastal mountains.

Elevation range: 730. Ataenius communis Hinton, 1936

Distribution: Mexico, Guatemala, El Salvador, Costa Rica, Panama, Colombia, Venezuela, and Trinidad & Tobago (^{Schoolmeesters, 2022}).

Locality records: Santa Ana: Parque Nacional Montecristo [probably San José Ingenio] (^{Stebnicka, 2001: 267}); La Libertad: Los Chorros (^{Stebnicka, 2001: 267}); Volcán de San Salvador (El Boquerón) (^{Stebnicka, 2001: 267}) (Fig. 6).

Years of collection: no data.

Months of collection: no data.

Topographic zone: coastal mountains, northern mountains.

Elevation range: 730-1,845 m. Ataenius complicatus Harold, 1869

Distribution: Mexico, Guatemala, Belize, El Salvador, Honduras, Nicaragua, Costa Rica, Panama, Colombia, Curaçao, Venezuela, Brazil, Peru, Bolivia, Paraguay, and Argentina (^{Schoolmeesters, 2022}).

Locality records: La Libertad: no specific locality (^{Stebnicka, 2006: 94}) (Fig. 6).

Years of collection: no data.

Months of collection: no data.

Topographic zone: no data.

Elevation range: no data. Ataenius cribrithorax Bates, 1887

Distribution: Mexico, El Salvador, Honduras, Costa Rica, Panama, Cuba, Jamaica, and US Virgin Islands (^{Schoolmeesters, 2022}).

Locality records: La Libertad: Santa Tecla (^{Stebnicka, 2001: 264}) (Fig. 6).

Years of collection: no data.

Months of collection: no data.

Topographic zone: coastal mountains.

Elevation range: 900 m. Ataenius glabriventris Schmidt, 1911

Distribution: Mexico, Guatemala, El Salvador, Honduras, Costa Rica, Panama, and Venezuela (^{Schoolmeesters, 2022}).

Locality records: La Libertad: no specific locality (^{Stebnicka & Lago, 2005: 72}) (Fig 6).

Years of collection: 1971.

Months of collection: May.

Topographic zone: no data.

Elevation range: no data.

Comments: the collection site is reported simply as “S Area” (^{Stebnicka & Lago, 2005}). It likely refers to an area near or in the city of La Libertad because of the date of collection: 11-V-1971 or dates near (13-V-1971). The collector of the specimen, H. Howden, collected other specimens around that area (^{Ratcliffe, 1978}; ^{Kohlmann, 1996}; ^{González-Alvarado & Vaz-de-Mello, 2014}; ^{Darling & Génier, 2018}).

Ataenius gracilis (Melsheimer, 1845)

Distribution: Canada, USA, Mexico, El Salvador, Nicaragua, Panama, Colombia, Ecuador (Galápagos Islands), Peru, Chile, Argentina, Cuba, Jamaica, Haiti, Dominican Republic, Puerto Rico, US Virgin Islands, St. Kitts & Nevis, Guadeloupe, Martinique, St. Vincent & the Grenadines, Barbados, and Grenada (^{Schoolmeesters, 2022}).

Locality records: no specific department: no specific locality (^{Berry, 1959b: 9}).

Years of collection: no data.

Months of collection: no data.

Topographic zone: no data.

Elevation range: no data.

Ataenius imbricatus (Melsheimer, 1844)

Distribution: Canada, USA, Mexico, Guatemala, Belize, El Salvador, Honduras, Nicaragua, Costa Rica, Panama, Colombia, Venezuela, Trinidad & Tobago, Guyana, Suriname, Brazil, Paraguay, Argentina, Bahamas, Cuba, Dominican Republic, Puerto Rico, US Virgin Islands, Guadeloupe, and Barbados (^{Schoolmeesters, 2022}).

Locality records: San Vicente: no specific locality (^{Stebnicka, 2003: 225}) (Fig 7).

Years of collection: no data.

Months of collection: no data.

Topographic zone: no data.

Elevation range: no data.

Ataenius liogaster Bates, 1887

Distribution: Mexico, Guatemala, El Salvador, Honduras, Nicaragua, Costa Rica, Panama, Venezuela, Jamaica, Dominican Republic, Puerto Rico, US Virgin Islands, Antigua & Barbuda, Guadeloupe, St. Lucia, St. Vincent & the Grenadines, Barbados, Grenada, Galápagos Islands. Australia: Christmas Island. Asia: Thailand, Vietnam, Philippines, Micronesia, and Indonesia (^{Stebnicka & Lago, 2005}; ^{Schoolmeesters, 2022}).

Locality records: La Libertad: Santa Tecla (^{Stebnicka & Lago, 2005: 61}) (Fig. 7). Years of collection: no data. Months of collection: no data. Topographic zone: coastal mountains. Elevation range: 900 m.

Ataenius nugator Harold, 1880

Distribution: Mexico, Guatemala, El Salvador, Honduras, Nicaragua, Costa Rica, Panama, Colombia, Venezuela, Ecuador, Brazil, Peru, Bolivia, Jamaica, Dominican Republic, US Virgin Islands, and Martinique (^{Schoolmeesters, 2022}).

Locality records: La Libertad: Los Chorros (^{Stebnicka, 2001: 263}) (Fig. 7).

Years of collection: no data.

Months of collection: no data.

Topographic zone: coastal mountains.

Elevation range: 730 m. Ataenius perforatus Harold, 1867

Distribution: Mexico, Guatemala, El Salvador, Honduras, Costa Rica, Panama, Colombia, Venezuela, and Brazil (^{Schoolmeesters, 2022}).

Locality records: La Libertad: Volcán de San Salvador (El Boquerón) (^{Stebnicka, 2001: 275}) (Fig. 7).

Years of collection: no data.

Months of collection: no data.

Topographic zone: coastal mountains.

Elevation range: 1,845 m. Ataenius platensis (Blanchard, 1846)

Distribution: Mexico, Guatemala, Belize, El Salvador, Honduras, Costa Rica, Panama, Colombia, Venezuela, Ecuador, Brazil, Peru, Bolivia, Chile, Paraguay, Argentina, Uruguay, and Guadeloupe (^{Schoolmeesters, 2022}).

Locality records: La Libertad: Los Chorros (^{Stebnicka, 2005: 126}) (Fig. 7).

Years of collection: 1980.

Months of collection: January.

Topographic zone: coastal mountains.

Elevation range: 730 m.

Comments: collection data from specimen in RDCC. Ataenius scalptifrons Bates, 1887

Distribution: Mexico, Guatemala, Belize, El Salvador, Honduras, and Costa Rica (^{Schoolmeesters, 2022}).

Locality records: no specific department: no specific locality (^{Galante et al., 2003: 293}).

Years of collection: no data.

Months of collection: no data.

Topographic zone: no data.

Elevation range: no data. Ataenius sculptor Harold, 1868

Distribution: USA, Mexico, Guatemala, El Salvador, Honduras, and Colombia (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Cara Sucia (^{Pablo-Cea et al., 2017: 528}); La Libertad: no specific locality (Stebnicka, 2006: 94) (Fig. 7).

Years of collection: 2016.

Months of collection: July.

Topographic zone: coastal plain.

Elevation range: 5 m. Ataenius texanus Harold, 1874

Distribution: USA, Mexico, Guatemala, El Salvador, and Costa Rica (^{Stebnicka, 2007}).

Locality records: no specific department: no specific locality (^{Stebnicka, 2007: 67}).

Years of collection: no data.

Months of collection: no data.

Topographic zone: no data.

Elevation range: no data. Ataenius usingeri Hinton, 1937

Distribution: Mexico, Guatemala, Belize, El Salvador, Honduras, Costa Rica, and Panama (^{Schoolmeesters, 2022}).

Locality records: La Libertad: La Libertad (^{Stebnicka, 2005: 133}); Santa Tecla (^{Stebnicka, 2005: 133}) (Fig. 7).

Years of collection: 1971

Months of collection: May and June.

Topographic zone: coastal plain, coastal mountains.

Elevation range: 70-900 m.

Odontolytes Koshantschikov, 1916 Odontolytes capitosus (Harold, 1867)

Distribution: Mexico, Guatemala, El Salvador, Nicaragua, Costa Rica, and Panama (^{Stebnicka, 2002}).

Locality records: La Libertad: Los Chorros (^{Stebnicka, 2002: 768}) (Fig. 7).

Years of collection: no data.

Months of collection: no data.

Topographic zone: coastal mountains.

Elevation range: 730 m.

Cartwrightia Islas, 1958 Cartwrightia islasi Cartwright, 1967

Distribution: Mexico, Guatemala, and El Salvador (^{Schoolmeesters, 2022}).

Locality records: San Salvador: San Salvador (^{Cartwright, 1967: 4}) (Fig. 7).

Years of collection: 1957.

Months of collection: May.

Topographic zone: interior valley.

Elevation range: 685 m.

Comments: the species was described from specimens collected at lights in San Salvador (^{Cartwright, 1967}).

Haroldiataenius Chalumeau, 1981 Haroldiataenius limbatus (Bates, 1887)

Distribution: Mexico, Guatemala, El Salvador, and Honduras (^{Stebnicka & Skelley 2007}; ^{Schoolmeesters, 2022}).

Locality records: San Salvador: San Salvador (^{Stebnicka & Skelley, 2007: 11}) (Fig. 7).

Years of collection: 1958.

Months of collection: June.

Topographic zone: interior valley.

Elevation range: 685 m.

Comments: specimens were captured in the detritus from a nest of Atta sp. (Hymenoptera: Formicidae) (^{Stebnicka & Skelley, 2007: 11}).

Tribe Psammodiini Mulsant, 1842 Trichiorhyssemus Clouët, 1901 Trichiorhyssemus cristatellus (Bates, 1887)

Distribution: Mexico, Guatemala, El Salvador, and Nicaragua (^{Schoolmeesters, 2022}).

Locality records: Sonsonate: no specific locality (^{Gordon & Cartwright, 1980: 20}) (Fig. 8).

Years of collection: no data.

Months of collection: no data.

Topographic zone: no data.

Elevation range: no data.

Subfamily Scarabaeinae Latreille, 1802 Tribe Ateuchini Perty, 1830 Ateuchus Weber, 1801

Ateuchus (Ateuchus) colossus^{Moctezuma, Sánchez-Huerta & Halffter, 2018}

Distribution: Mexico and El Salvador (^{Moctezuma et al., 2018}, M. Cupello, personal communication).

Locality records: Santa Ana: Parque Nacional Montecristo (Bosque Nebuloso, La Torre [Camino hacia la Hacienda Montecristo], Los Planes) (^{Kohlmann, 2000: 244}) (Fig. 8).

Years of collection: 1971.

Months of collection: May.

Topographic zone: northern mountains.

Elevation range: 1,450-2,360 m.

Comments: the specimens ^{Kohlmann (2000)} identified as Ateuchus chrysopyge from El Salvador are A. colossus. Ateuchus chrysopyge is a junior synonym of A. illaesus and does not occur on the Pacific side of Central America (M. Cupello, personal communication).

Ateuchus (Ateuchus) rodriguezi (Preudhomme de Borre, 1886)

Distribution: Mexico, Guatemala, El Salvador, Honduras, Nicaragua, and Costa Rica (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (^{Pablo-Cea, 2014: 46}, ^{Pablo-Cea et al., 2020: 6}; ^{Pablo-Cea, 2021: 20}; ^{Pablo-Cea, Deloya, MacGregor-Fors, Navarrete-Heredia, & Espinosa-de los Monteros, 2022: 378}); La Libertad: Los Chorros (^{Kohlmann, 1996: 187}); Parque Nacional Walter Thilo Deininger (^{Fuentes, 1998: 29}; ^{Horgan, 2008: 2968}); No specific locality (MZSP); San Salvador: Apopa (Josué Chávez, personal communication); No specific locality (CMNC, CNCI); Cuscatlán: Cojutepeque (MZSP); La Paz: Comalapa (La Providencia) (^{Horgan, 2002: 30}); San Vicente: Volcán de San Vicente-Chinchontepec (Finca La Paz) (MZSP); No specific locality (DZUP); San Miguel: San Miguel (EMEC); La Unión: La Unión (^{Berry & Salazar-Vaquero, 1957: 43}) (Fig. 8).

Years of collection: 1959-1962, 1971, 1995-1998, 2013, 2018.

Months of collection: April-July, September, and October.

Topographic zone: interior valley, coastal mountains, coastal plain.

Elevation range: 30-2,000 m.

Comments: reported as Ateuchus ampliatus Bates by ^{Berry and Salazar-Vaquero (1957)}. The species is rare in Parque Nacional El Imposible (^{Pablo-Cea, 2014}, ²⁰²¹). Specimens were captured with baited pitfall traps in Parque Nacional Walter Thilo Deininger (cow dung, beef carrion, and fruit) (^{Horgan, 2008}), Comalapa (cow and horse dung) (^{Horgan, 2001}), Cara Sucia (^{Fuentes, 2009}; ^{Pablo-Cea et al., 2017}), and Parque Nacional El Imposible (human dung) (^{Pablo-Cea, 2014}, ²⁰²¹).

Scatimus Erichson, 1847

Scatimus ovatus Harold, 1862

Distribution: Mexico, Guatemala, Belize, El Salvador, Nicaragua, Costa Rica, Panama, and Colombia (^{Schoolmeesters, 2022}).

Locality records: no specific department: no specific locality (^{Howden & Young, 1981: 50}).

Years of collection: no data.

Months of collection: no data.

Topographic zone: no data.

Elevation range: no data.

Uroxys Westwood, 1842

Uroxys boneti Pereira & Halffter, 1961

Distribution: Mexico, Guatemala, El Salvador, Nicaragua, Costa Rica, Panama, and Colombia (^{Schoolmeesters, 2022}).

Locality records: no specific department: no specific locality (^{Solís & Kohlmann, 2013: 293}).

Years of collection: no data.

Months of collection: no data.

Topographic zone: no data.

Elevation range: no data.

Uroxys deavilai Delgado & Kohlmann, 2007 Distribution: Mexico, Guatemala, El Salvador,

Nicaragua, and Costa Rica (^{Schoolmeesters, 2022}). Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (^{Pablo-Cea, 2014: 46}; ^{Pablo-Cea et al., 2016: 220}, ^{2020: 6}) (Fig. 8).

Years of collection: 2013.

Months of collection: June-November.

Topographic zone: coastal mountains.

Elevation range: 780 m.

Comments: specimens were captured in pitfall traps baited with human dung and chicken carrion (^{Pablo-Cea, 2014}).

Uroxys microcularis Howden & Young, 1981 Distribution: Mexico, El Salvador, Costa Rica, and

Panama (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (^{Pablo-Cea, 2014: 46}, ^{Pablo-Cea et al., 2016: 220}; ^{Pablo-Cea et al., 2020: 6}) (Fig. 8).

Years of collection: 2013.

Months of collection: June-November.

Topographic zone: coastal mountains.

Elevation range: 780 m.

Comments: specimens were captured in pitfall traps baited with human dung and chicken carrion (^{Pablo-Cea, 2014}).

Tribe Coprini Leach, 1815

Canthidium Erichson, 1847

Canthidium (Canthidium) laetum Harold, 1867 Distribution: Mexico, Guatemala, El Salvador,

Honduras, Nicaragua, and Costa Rica (^{Schoolmeesters, 2022}).

Locality records: La Paz: El Rosario (^{Solís & Kohlmann, 2004: 49}; ^{Kohlmann & Solís, 2006: 255}); La Unión: La Unión (^{Solís & Kohlmann, 2004: 49}; ^{Kohlmann & Solís, 2006: 255}) (Fig. 8). Years of collection: 1958. Months of collection: May. Topographic zone: coastal plain. Elevation range: 30-85 m. Comments: captured under dung.

Canthidium (Canthidium) pseudopuncticolle^{Solís & Kohlmann, 2004}

Distribution: USA, Mexico, El Salvador, and Costa Rica (^{Solís & Kohlmann, 2004}, ^{Pablo-Cea et al., 2016}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (^{Pablo-Cea, 2014: 46}, ^{Pablo-Cea et al., 2016: 219-220}, ^{2020: 6}) (Fig. 8).

Years of collection: 2013.

Months of collection: June-November.

Topographic zone: coastal mountains.

Elevation range: 780 m.

Comments: reported as Canthidium pseudopunticolle [sic] by ^{Pablo-Cea (2014)} and ^{Pablo-Cea et al. (2016}, ²⁰²⁰⁾. Specimens were captured in pitfall traps baited with human dung and chicken carrion (^{Pablo-Cea, 2014}).

Copris Geoffroy, 1762

Copris (Copris) aspericollis Gillet, 1910

Distribution: Guatemala, El Salvador, and Costa Rica (^{Schoolmeesters, 2022}).

Locality records: Santa Ana: Parque Nacional Montecristo (Los Planes) (^{Kohlmann et al., 2003: 13}) (Fig. 8).

Years of collection: 2002.

Months of collection: June.

Topographic zone: northern mountains.

Elevation range: 1,890 m.

Copris (Copris) boucardi Harold, 1869

Distribution: Mexico, Guatemala, and El Salvador (^{Schoolmeesters, 2022}).

Locality records: Santa Ana: Parque Nacional Montecristo (Bosque Nebuloso) (FMNH); no specific department: no specific locality (^{Deloya et al., 2014: 73}) (Fig. 8).

Years of collection: 1971.

Months of collection: May.

Topographic zone: northern mountains.

Elevation range: 2,360 m.

Copris (Copris) costaricensis Gahan, 1894

Distribution: Mexico, El Salvador, Costa Rica, and Panama (^{Fuentes, 2009}; ^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (^{Pablo-Cea, 2014: 46}; ^{Pablo-Cea et al., 2020: 6}; ^{Pablo-Cea, Deloya, MacGregor-Fors, Navarrete-Heredia, & Espinosa-de los Monteros, 2022: 378}); La Libertad: Parque El Bicentenario (^{Pablo-Cea, 2021: 20}; ^{Pablo-Cea, Deloya, MacGregor-Fors, & Navarrete-Heredia, 2022: 277}; ^{Pablo-Cea, Deloya, MacGregor-Fors, Navarrete-Heredia et al., 2022: 378}); San Salvador: Tonacatepeque (^{Fuentes, 2009: 22}) (Fig. 8).

Years of collection: 2007, 2013, 2018.

Months of collection: June-November.

Topographic zone: interior valley, coastal mountains.

Elevation range: 625-870 m.

Comments: specimens were captured in pitfall traps baited with human dung and chicken carrion in Parque Nacional El Imposible (^{Pablo-Cea, 2014}), cattle dung in Tonacatepeque (^{Fuentes, 2009}), and squid carrion in Parque El Bicentenario (^{Pablo-Cea, 2021}).

Copris (Copris) laeviceps Harold, 1869

Distribution: Mexico, Guatemala, Belize, El Salvador, Honduras, Nicaragua, Costa Rica, and Colombia (^{Horgan, 2001}; ^{Schoolmeesters, 2022}).

Locality records: Morazán: Jocoaitique (El Rodeo) (^{Horgan, 2001: 106}) (Fig. 9).

Years of collection: 1997.

Months of collection: May and August.

Topographic zone: northern mountains.

Elevation range: 655 m.

Comments: specimens were captured in pitfall traps baited with cow dung.

Copris (Copris) lugubris Boheman, 1858 Distribution: Mexico, Guatemala, Belize, El

Salvador, Honduras, Nicaragua, Costa Rica, and Panama (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Cara Sucia (^{Pablo-Cea et al., 2017: 528}); Parque Nacional El Imposible (La Fincona) (MUHNES); (San Benito) (^{Pablo-Cea, 2014: 46}; ^{Darling & Génier, 2018: 21}; ^{Pablo-Cea et al., 2020: 6}); Santa Ana: Metapán (^{Darling & Génier, 2018: 21}); Santa Ana (^{Darling & Génier, 2018: 21}); Chalatenango: Chalatenango (^{Darling & Génier, 2018: 21}); La Palma (^{Darling & Génier, 2018: 21}); La Libertad: La Libertad (^{Darling & Génier, 2018: 21}); Los Chorros (^{Darling & Génier, 2018: 21}); Parque Nacional Walter Thilo Deininger (^{Fuentes, 1998: 29}; ^{Horgan, 2008: 2968}); Santa Tecla (^{Darling & Génier, 2018: 21}); San Salvador: Aguilares (^{Darling & Génier, 2018: 21}); San Salvador (^{Darling & Génier, 2018: 21}); (Universidad de El Salvador) (Josué Chávez and Melissa Oviedo, personal communication); Tonacatepeque (^{Fuentes, 2009: 22}); La Paz: Comalapa (MUHNES); (La Providencia) (^{Fuentes, 1998: 29}; ^{Horgan, 2001: 106}; ^{Horgan, 2002: 30}; ^{Horgan, 2008: 2969}); Cuscatlán: Área Natural Protegida Colima (Cerrón Grande) (J. Pablo, personal observation); No specific locality (^{Darling & Génier, 2018: 21}); Cabañas: Chorrera del Guayabo (^{Darling & Génier, 2018: 21}); Usulután: Cerro El Tigre (MUHNES); San Miguel: Laguna El Jocotal (MUHNES); Morazán: Arambala (Río Sapo) (MUHNES); Cerro Cacahuatique (MUHNES); Cerro Perquín (^{Fuentes, 1998: 29}; ^{Horgan, 2001: 106}); Jocoaitique (El Rodeo) (Fuentes, 1998: 29; Horgan, 2001: 106); La Unión: Volcán de Conchagua (^{Darling & Génier, 2018: 21}); no specific department: no specific locality (^{Howden & Young, 1981: 132}) (Fig. 9).

Years of collection: 1958-1960, 1971, 1979, 1995-2001, 2007, 2012-2013, 2016.

Months of collection: February-December.

Topographic zone: northern mountains, interior valley, coastal mountains, coastal plain.

Elevation range: 5-1,580 m.

Comments: specimens were captured in baited pitfall traps in Parque Nacional Walter Thilo Deininger (cow dung, beef carrion, fruit) (^{Horgan, 2008}), Comalapa (beef carrion, cow and horse dung) (^{Horgan, 2001}, ²⁰⁰⁸), Jocoaitique and Cerro Perquín (cow and horse dung) (^{Horgan, 2001}), Cara Sucia and Tonacatepeque (cow dung) (^{Fuentes, 2009}; ^{Pablo-Cea et al., 2017}), and Parque Nacional El Imposible (human dung) (^{Pablo-Cea, 2014}). Copris (Copris) matthewsi pacificus^{Delgado & Kohlmann, 2001}

Distribution: Mexico, Guatemala, and El Salvador (^{Schoolmeesters, 2022}).

Locality records: Santa Ana: Parque Nacional Montecristo (Bosque Nebuloso) (^{Delgado & Kohlmann, 2001: 350}); Chalatenango: Cerro El Pital (^{Kohlmann et al., 2003: 15}) (Fig. 9).

Years of collection: 1971, 2002.

Months of collection: May, June, and September.

Topographic zone: northern mountains.

Elevation range: 2,360-2,660 m.

Dichotomius Hope, 1838

Dichotomius (Dichotomius) annae^{Kohlmann & Solís, 1997}

Distribution: Mexico, Guatemala, El Salvador, Honduras, Nicaragua, Costa Rica, and Panama (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (^{Pablo-Cea, 2014: 46}; ^{Pablo-Cea et al., 2020: 6}); Santa Ana: Parque Nacional Montecristo (Los Planes) (RDCC); Sonsonate: Sonsonate (^{Kohlmann & Solís, 1997: 356}); La Libertad: La Libertad (^{Kohlmann & Solís, 1997: 356}); Parque El Bicentenario (^{Pablo-Cea, 2021: 20}; ^{Pablo-Cea, Deloya, MacGregor-Fors, & Navarrete-Heredia, 2022: 277}; ^{Pablo-Cea, Deloya, MacGregor-Fors, Navarrete-Heredia et al., 2022: 378}); Parque Nacional Walter Thilo Deininger (^{Horgan, 2008: 2968}); San Andrés (MUHNES); San Salvador: Parque Saburo Hirao (MUHNES); San Salvador (^{Kohlmann & Solís, 1997: 356}); Tonacatepeque (^{Fuentes, 2009: 22}); La Paz: Comalapa (Aeropuerto) (MUHNES); (La Providencia) (^{Horgan, 2008: 2969}); San Miguel: Laguna El Jocotal (MUHNES); Morazán: Arambala (Río Sapo) (MUHNES) (Fig. 9).

Years of collection: 1905, 1920, 1953, 1958, 1979, 1998-2001, 2013, 2018-2019.

Months of collection: April-September, and November.

Topographic zone: northern mountains, interior valley, coastal mountains, coastal plain.

Elevation range: 35-1,890 m.

Comments: specimens were captured in baited pitfall traps in Parque Nacional Walter Thilo Deininger and Comalapa (beef carrion and cow dung) (^{Horgan, 2008}), Parque Nacional El Imposible and Parque El Bicentenario (human dung) (^{Pablo-Cea, 2014}; ^{Pablo-Cea, 2021}), and Jocoaitique (cow dung) (^{Horgan, 2001}).

Dichotomius (Selenocopris) centralis (Harold, 1869) Distribution: Guatemala, Belize, El Salvador, Nicaragua, and Costa Rica (^{Schoolmeesters, 2022}). Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (^{Pablo-Cea, 2014: 46}; ^{Pablo-Cea et al., 2020: 6}); Chalatenango: Comalapa (^{Kohlmann & Solís, 1997: 358}); La Libertad: Parque El Bicentenarion (^{Pablo-Cea, 2021: 20}; ^{Pablo-Cea, Deloya, MacGregor-Fors, & Navarrete-Heredia, 2022: 277}; ^{Pablo-Cea, Deloya, MacGregor-Fors, Navarrete-Heredia et al., 2022: 378}); Parque Nacional Walter Thilo Deininger (^{Fuentes, 1998: 29}; ^{Horgan, 2008: 2968}); San Salvador: San Salvador (^{Berry & Salazar-Vaquero, 1957: 47}); (Universidad de El Salvador) (Josué Chávez & Melissa Oviedo, personal communication); Tonacatepeque (^{Fuentes, 2009: 22}); La Paz: Comalapa (La Providencia) (^{Fuentes, 1998: 29}; ^{Horgan, 2001: 106}, ^{2002: 30}, ^{2008: 2969}); Morazán: Cerro Perquín (^{Fuentes, 1998: 29}; ^{Horgan, 2001: 106}); Jocoaitique (El Rodeo) (^{Fuentes, 1998: 29}; ^{Horgan, 2001: 106}) (Fig. 9).

Years of collection: 1976, 1995-1998, 2008, 2013, 2018-2019.

Months of collection: April-November.

Topographic zone: northern mountains, interior valley, coastal mountains, coastal plain.

Elevation range: 40-1,205 m.

Comments: reported as Pinotus centralis by ^{Berry and Salazar-Vaquero (1957)}. Specimens were captured in baited pitfall traps in Parque Nacional Walter Thilo Deininger (beef carrion, cow and horse dung, fruit) (^{Fuentes, 1998}, ^{Horgan, 2008}), Comalapa (La Providencia) (beef carrion, cow and horse dung) (^{Horgan, 2001}, ²⁰⁰⁸), Parque El Bicentenario (squid carrion, human dung, decaying fruit) (^{Pablo-Cea, 2021}), Parque Nacional El Imposible (chicken carrion, human dung) (^{Pablo-Cea, 2014}), Jocoaitique (El Rodeo) and Cerro Perquín (cow and horse dung) (^{Horgan, 2001}), and Tonacatepeque (cow dung) (^{Fuentes, 2009}).

Dichotomius (Selenocopris) yucatanus (Bates, 1887) Distribution: Mexico, El Salvador, Honduras,

Nicaragua, Costa Rica, and Colombia (^{Fuentes, 1998}; ^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (^{Pablo-Cea, 2014: 46}; ^{Pablo-Cea et al., 2020: 6}); Chalatenango: San Ignacio (MUHNES); La Libertad: Parque El Bicentenario (^{Pablo-Cea, 2021: 20}; ^{Pablo-Cea, Deloya, MacGregor-Fors, & Navarrete-Heredia, 2022: 277}; ^{Pablo-Cea, Deloya, MacGregor-Fors, Navarrete-Heredia et al., 2022: 378}); Parque Nacional Walter Thilo Deininger (^{Fuentes, 1998: 29}; ^{Horgan, 2008: 2968}); Cabañas: Cinquera (RDCC) (Fig. 9).

Years of collection: 1979, 1995-1998, 2001, 2013, 2018-2019.

Months of collection: April-November.

Topographic zone: northern mountains, interior valley, coastal mountains, coastal plain.

Elevation range: 135-1,060 m.

Comments: specimens were captured in baited pitfall traps in Parque El Bicentenario (squid carrion, human dung, decaying fruit) (^{Pablo-Cea, 2021}), Parque Nacional Walter Thilo Deininger (beef carrion, cow and horse dung) (^{Fuentes, 1998}; ^{Horgan, 2008}), and Parque Nacional El Imposible (chicken carrion, human dung) (^{Pablo-Cea, 2014}).

Tribe Deltochilini Lacordaire, 1856 Agamopus Bates, 1887 Agamopus lampros Bates, 1887

Distribution: El Salvador, Nicaragua, Costa Rica, Panama, and Colombia (^{Fuentes, 1998}; ^{Schoolmeesters, 2022}).

Locality records: La Libertad: Parque Nacional Walter

Thilo Deininger (^{Fuentes, 1998: 29}; ^{Horgan, 2008: 2968}); no specific department: no specific locality (^{Howden & Young, 1981: 41}) (Fig. 9).

Years of collection: 1995-1997.

Months of collection: no data.

Topographic zone: coastal plain.

Elevation range: 135 m.

Comments: specimens were captured in pitfall traps baited with beef carrion, cow dung, and fruit (^{Horgan, 2008}).

Canthon Hoffmannsegg, 1817

Canthon (Canthon) cyanellus LeConte, 1859 Distribution: USA, Mexico, Guatemala, El Salvador, Nicaragua, Costa Rica, Panama, Colombia, and Venezuela (^{Fuentes, 1998}; ^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (^{Pablo-Cea, 2014: 46}; ^{Pablo-Cea et al., 2020: 6}); La Libertad: Parque Nacional Walter Thilo Deininger (^{Horgan, 2008: 2968}); San Salvador: Tonacatepeque (^{Fuentes, 2009: 22}); Morazán: Jocoaitique (El Rodeo) (^{Fuentes, 1998}; ^{Horgan, 2001: 106}) (Fig. 9).

Years of collection: 1995-1997.

Months of collection: July and August.

Topographic zone: northern mountains, interior valley, coastal mountains, coastal plain.

Elevation range: 135-780 m.

Comments: reported as Canthon cyanellis [sic] by ^{Fuentes (1998)}. Specimens were captured in baited pitfall traps in Parque Nacional El Imposible (chicken carrion) (^{Pablo-Cea, 2014}), Tonacatepeque and Jocoaitique (cow dung) (^{Fuentes, 1998}, ^{Horgan, 2008}), and Parque Nacional Walter Thilo Deininger (cow dung, beef carrion, fruit) (^{Horgan, 2008}).

Canthon (Canthon) deyrollei Harold, 1868

Distribution: Guatemala, Belize, El Salvador, Nicaragua, Costa Rica, Panama, and Colombia (Solís & Kohlmann, 2002; ^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (MUHNES, RDCC); San Vicente: Apastepeque (Poza Azul) (MUHNES, RDCC); no specific department: no specific locality (^{Berry, 1959b: 15}) (Fig. 9).

Years of collection: 1979.

Months of collection: June.

Topographic zone: interior valley, coastal mountains.

Elevation range: 500-780 m.

Comments: specimens from Parque Nacional El Imposible were collected in horse dung (R. D. Cave, personal observation).

Canthon (Canthon) indigaceus chevrolati Harold, 1868 Distribution: Mexico, Guatemala, El Salvador, Honduras, Nicaragua, Costa Rica, Panama, and Colombia (Solís & Kohlmann, 2002; ^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Francisco Menéndez) (MUHNES, RDCC); La Libertad: Parque Nacional Walter Thilo Deininger (^{Fuentes, 1998: 29}; ^{Horgan, 2008: 2968}); San Salvador: Tonacatepeque (^{Fuentes, 2009: 22}); La Paz: Comalapa (MUHNES); (La Providencia) (^{Horgan, 2001: 106}, ^{2002: 30}, ^{2008: 2969}); San Vicente: Apastepeque (Poza Azul) (MUHNES, RDCC); Tecoluca (^{Berry & Salazar-Vaquero, 1957: 44}) (Fig. 9).

Years of collection: 1979, 1995-1997, 2001.

Months of collection: May-August.

Topographic zone: interior valley, coastal plain.

Elevation range: 40-625 m.

Comments: reported as Canthon chevrolati by Berry and ^{Salazar-Vaquero (1957)}. Specimens were captured in baited pitfall traps in Parque Nacional Walter Thilo Deininger (cow and horse dung) (^{Fuentes, 2008}; ^{Horgan, 2008}) and Tonacatepeque and Comalapa (cow dung) (^{Horgan, 2008}; ^{Fuentes, 2009}).

Canthon (Canthon) morsei Howden, 1966

Distribution: Mexico, El Salvador, Nicaragua, Costa Rica, Panama, Colombia, and Ecuador (Solís & Kohlmann, 2002; ^{Schoolmeesters, 2022}).

Locality records: no specific department: no specific locality (^{Howden & Young, 1981: 29}).

Years of collection: no data.

Months of collection: no data.

Topographic zone: no data.

Elevation range: no data.

Canthon (Glaphyrocanthon) championi Bates, 1887 Distribution: Mexico, Guatemala, El Salvador, Costa Rica, and Panama (^{Rivera-Cervantes & Halffter 1999}; ^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Francisco Menéndez) (MUHNES); La Libertad: Parque Nacional Walter Thilo Deininger (MUHNES); no specific locality (^{Rivera-Cervantes & Halffter, 1999: 86}) (Fig. 10).

Years of collection: 1971, 1979, 1995, 2002.

Months of collection: May and September.

Topographic zone: coastal plain.

Elevation range: 135-250 m.

Canthon (Glaphyrocanthon) femoralis (Chevrolat, 1834) Distribution: Mexico, Guatemala, Belize, El Salvador, Honduras, Nicaragua, Costa Rica, Panama, Colombia, Ecuador, and Bolivia (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (^{Pablo-Cea, 2014: 46}; ^{Pablo-Cea et al., 2020: 6}); La Libertad: Los Chorros (MUHNES) (Fig. 10); no specific department: no specific locality (^{Morón, 2003: 32}).

Years of collection: 1979, 2013.

Months of collection: June-November.

Topographic zone: coastal mountains.

Elevation range: 730-780 m.

Canthon (Glaphyrocanthon) meridionalis Martínez, Halffter, & Halffter, 1964

Distribution: El Salvador, Honduras, Nicaragua, Costa Rica, and Panama (^{Horgan, 2007}; ^{Schoolmeesters, 2022}).

Locality records: La Libertad: Parque Nacional Walter Thilo Deininger (^{Fuentes, 2008: 29}; ^{Horgan, 2008: 2968}) (Fig. 10).

Years of collection: 1995-1998.

Months of collection: no data.

Topographic zone: coastal plain.

Elevation range: 135 m.

Comments: reported as Canthon viridis meridionalis by ^{Horgan (2007}, ²⁰⁰⁸⁾. Specimens were captured in pitfall traps baited with beef carrion, cow and horse dung, and fruit (^{Fuentes, 1998}; ^{Horgan, 2008}).

Deltochilum Eschscholtz, 1822

Deltochilum (Calhyboma) mexicanum Burmeister, 1848 Distribution: Mexico, Guatemala, Belize, El Salvador, Honduras, Nicaragua, Costa Rica, Panama, Colombia, Ecuador, Peru, and Bolivia (^{Schoolmeesters, 2022}).

Locality records: no specific department: no specific locality (^{Howden & Young, 1981: 38}).

Years of collection: no data.

Months of collection: no data.

Topographic zone: no data.

Elevation range: no data.

Deltochilum (Deltochilum) scabriusculum Bates, 1887 Distribution: USA, Mexico, Belize, Guatemala, El Salvador, Nicaragua, and Costa Rica (^{Schoolmeesters, 2022}).

Locality records: San Salvador: Tonacatepeque (^{Fuentes, 2009: 22}) (Fig. 10).

Years of collection: 1997.

Months of collection: no data.

Topographic zone: interior valley.

Elevation range: 625 m.

Comments: reported as Deltochilium [sic] scabriusculum by ^{Fuentes (2009)}.

Deltochilum (Hybomidium) lobipes Bates, 1887 Distribution: Mexico, Guatemala, Belize, El Salvador, Honduras, Nicaragua, Costa Rica, and Colombia (^{Schoolmeesters, 2022}).

Locality records: La Libertad: La Libertad (^{González-Alvarado & Vaz-de-Mello, 2014: 445}); Parque Nacional Walter Thilo Deininger (^{Horgan, 2008: 2968}); La Paz: Comalapa (La Providencia) (^{Horgan, 2002: 30}) (Fig. 10).

Years of collection: 1971, 1995-1998.

Months of collection: May.

Topographic zone: coastal plain.

Elevation range: 40-135 m.

Comments: specimens were captured in baited pitfall traps in Parque Nacional Walter Thilo Deininger (cow dung, beef carrion) and Comalapa (cow dung) (^{Horgan, 2002}, ²⁰⁰⁸).

Deltochilum (Hybomidium) sublaeve Bates, 1887 Distribution: USA, Mexico, Guatemala, Belize, El Salvador, Nicaragua, Costa Rica, Panama, and Ecuador (^{Howden & Young, 1981}; ^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (^{Pablo-Cea, 2014: 46}; ^{Pablo-Cea et al., 2020: 6}) (Fig. 10); no specific department: no specific locality (^{Howden & Young, 1981: 37}).

Years of collection: 2013.

Months of collection: June, and August-October.

Topographic zone: coastal mountains.

Elevation range: 780 m.

Comments: reported as Deltochilum (Hybomidium) gibbosum panamensis Howden by ^{Howden & Young (1981)}. Reported as D. gibbosum by ^{Pablo-Cea (2014)} and ^{Pablo-Cea et al. (2020)}.

Malagoniella Martínez, 1961

Malagoniella (Malagoniella) yucateca (Harold, 1863) Distribution: USA, Mexico, El Salvador, Nicaragua, and Costa Rica (^{Horgan, 2008}; ^{Schoolmeesters, 2022}). Locality records: La Libertad: Parque Nacional Walter

Thilo Deininger (^{Horgan, 2008: 2968}) (Fig. 10).

Years of collection: 1995-1997.

Months of collection: no data.

Topographic zone: coastal plain.

Elevation range: 135 m.

Comments: reported as Malagoniella astyanax (Olivier) by ^{Horgan (2008)}. Specimens were captured in pitfall traps baited with cow dung (^{Horgan, 2008}).

Pseudocanthon Bates, 1887 Pseudocanthon perplexus (LeConte, 1847)

Distribution: USA, Mexico, Guatemala, El Salvador, Honduras, Nicaragua, Costa Rica, Panama, Colombia, Venezuela, Brazil, and Grand Bahama Island (^{Schoolmeesters, 2022}).

Locality records: Usulután: Nancuchiname (MUHNES) (Fig. 10); no specific department: no specific locality (^{Howden & Young, 1981: 35}).

Years of collection: 1976.

Months of collection: May.

Topographic zone: coastal plain.

Elevation range: 15 m.

Comments:^{Horgan (2007)} reported Pseudocanthon sp. 1, which probably is P. perplexus.

Tribe Oniticellini Kolbe, 1905

Euoniticellus Janssens, 1953 Euoniticellus intermedius (Reiche, 1848)

Distribution: USA, Mexico, Guatemala, El Salvador, Honduras, Nicaragua, Costa Rica, and Panama; and Afrotropical and Palearctic regions (^{Solís et al., 2015}; ^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Cara Sucia (^{Pablo-Cea et al., 2017: 527}); La Libertad: Laguna Caldera (^{Pablo-Cea et al., 2017: 527}); Playa El Majahual (^{Pablo-Cea et al., 2017: 527}); San Salvador: Tonacatepeque (^{Fuentes, 2008: 32}; ^{Solís & Kohlmann, 2012: 13}; ^{Pablo-Cea et al., 2017: 529}); Cuscatlán: Área Natural Protegida Colima (Cerrón Grande) (^{Pablo-Cea et al., 2017: 527}); San Vicente: Eco-Parque Tehuacán (^{Pablo-Cea et al., 2017: 527}) (Fig. 11).

Years of collection: 2003-2008, 2012-2016.

Months of collection: July.

Topographic zone: interior valley, coastal plain.

Elevation range: 5-625 m.

Eurysternus Dalman, 1824 Eurysternus mexicanus Harold, 1869

Distribution: Mexico, Guatemala, Belize, El Salvador, Honduras, Nicaragua, Costa Rica, Panama, Trinidad and Tobago, Colombia, Venezuela, and Guyana (^{Horgan, 2001}; ^{Schoolmeesters, 2022}).

Locality records: Morazán: Cerro Perquín (^{Horgan, 2001: 106}) (Fig. 11).

Years of collection: no data.

Months of collection: no data.

Topographic zone: northern mountains.

Elevation range: 1,205 m.

Comments: specimens were captured in pitfall traps baited with cow dung (^{Horgan, 2001}).

Eurysternus magnus Laporte, 1840

Distribution: Mexico, Guatemala, Belize, El Salvador, Honduras, Nicaragua, Costa Rica, and Panama (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (^{Pablo-Cea, 2014: 46}; ^{Pablo-Cea et al., 2020: 6}); Santa Ana: Parque Nacional Cerro Verde (^{Génier, 2009: 191}) (Fig. 11).

Years of collection: 1971, 2013.

Months of collection: May, July-September, and November.

Topographic zone: coastal mountains.

Elevation range: 780-2,020 m.

Comments: captured in pitfall traps baited with cow dung.

Tribe Onthophagini Burmeister, 1846 Digitonthophagus Balthasar, 1959 Digitonthophagus gazella (Fabricius, 1787)

Distribution: USA, Mexico, Guatemala, El Salvador, Colombia, Brazil, Peru, Argentina, Cuba, Jamaica, Dominican Republic, and Guadeloupe; and Afrotropical region, Asia, and Australia (^{Noriega et al., 2015}; ^{Génier & Moretto 2017}; ^{Pablo-Cea et al., 2017}; ^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Cara Sucia (^{Pablo-Cea et al., 2017: 527}) (Fig. 11).

Years of collection: 2016.

Months of collection: July.

Topographic zone: coastal plain.

Elevation range: 5 m.

Onthophagus Latreille, 1802

Onthophagus (Onthophagus) batesi^{Howden & Cartwright, 1963}

Distribution: USA, Mexico, Guatemala, El Salvador, Honduras, Nicaragua, Costa Rica, Panama, Colombia, and Martinique (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (^{Pablo-Cea, 2014: 46}; ^{Pablo-Cea et al., 2020: 6}); Sonsonate: Sonsonate (^{Howden & Cartwright, 1963: 23}); Chalatenango: San Ignacio (RDCC); La Libertad: La Ceiba (^{Howden & Cartwright, 1963: 23}); Parque El Bicentenario (^{Pablo-Cea, 2021: 20}; ^{Pablo-Cea, Deloya, MacGregor-Fors, & Navarrete-Heredia, 2022: 277}; ^{Pablo-Cea, Deloya, MacGregor-Fors, Navarrete-Heredia et al., 2022: 378}); Parque Nacional Walter Thilo Deininger (^{Horgan, 2008: 2968}); Santa Tecla (^{Howden & Cartwright, 1963: 23}); San Salvador: Ilopango (^{Berry & Salazar-Vaquero, 1957: 46}); Mejicanos (J. Pablo, personal observation; Parque Saburo Hirao (MUHNES, RDCC); San Salvador (^{Howden & Cartwright, 1963: 23}); (Universidad de El Salvador) (Josué Chávez and Melissa Oviedo, personal communication); Tonacatepeque (^{Howden & Cartwright, 1963: 23}; ^{Fuentes, 2009: 22}); Cuscatlán: El Rosario (^{Berry & Salazar-Vaquero, 1957: 46}); La Paz: Comalapa (MUHNES); (La Providencia) (^{Horgan, 2001: 106}, ^{2002: 30}, ^{2008: 2969}); San Miguel: Ciudad Barrios (Cerro Cacahuatique) (^{Howden & Cartwright, 1963: 23}); Laguna Olomega (^{Howden & Cartwright, 1963: 23}); Morazán: Jocoaitique (El Rodeo) (^{Horgan, 2001: 106}); La Unión: La Unión (^{Howden & Cartwright, 1963: 23}; ^{Kohlmann & Solís, 2001: 175}; ^{Pulido-Herrera & Zunino, 2007: 96}); Volcán de Conchagua (^{Howden & Cartwright, 1963: 23}) (Fig. 11).

Years of collection: 1925, 1958, 1979, 1995-1998, 2013, 2018-2019.

Months of collection: All year, except March.

Topographic zone: northern mountains, interior valley, coastal mountains, coastal plain.

Elevation range: 30-1,405 m.

Comments: reported as O. incensus by Berry and ^{Salazar-Vaquero (1957)} (see O. incensus for more details). The species was described based on specimens from La Unión (^{Howden & Cartwright, 1963: 23}). Reported as Onthofagus batisi [sic] by ^{Fuentes (2009)}. Specimens were captured in baited pitfall traps in Comalapa (beef carrion, cow and horse dung) (^{Horgan, 2008}), Parque El Bicentenario (squid carrion, human dung, decaying fruit) (^{Pablo-Cea, 2021}), Parque Nacional Walter Thilo Deininger (beef carrion, cow dung) (^{Horgan, 2008}), Jocoaitique (cow and horse dung) (^{Horgan, 2001}), Parque Nacional El Imposible (chicken carrion, human dung) (^{Pablo-Cea, 2014}), and Tonacatepeque (cow dung) (^{Fuentes, 2009}).

Onthophagus (Onthophagus) belorhinus (Bates, 1887) Distribution: Mexico, Guatemala, El Salvador, Costa Rica, Colombia, and Ecuador (^{Schoolmeesters, 2022}). Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (^{Pablo-Cea, 2014: 46}; ^{Pablo-Cea et al., 2020: 6}); La Libertad: Parque El Bicentenario (^{Pablo-Cea, 2021: 20}; ^{Pablo-Cea, Deloya, MacGregor-Fors, & Navarrete-Heredia, 2022: 277}; ^{Pablo-Cea, Deloya, MacGregor-Fors, Navarrete-Heredia et al., 2022: 378}) (Fig. 11); no specific department: no specific locality (^{Howden & Young, 1981: 121}).

Years of collection: 2013, 2018-2019.

Months of collection: All year.

Topographic zone: coastal mountains.

Elevation range: 780-870 m.

Comments: specimens were captured in baited pitfall traps in Parque El Bicentenario (squid carrion, human dung, decaying fruit) (^{Pablo-Cea, 2021}) and Parque Nacional El Imposible (human dung, chicken carrion) (^{Pablo-Cea, 2014}).

Onthophagus (Onthophagus) championi Bates, 1887 Distribution: México, Guatemala, El Salvador, and Costa Rica (^{Fuentes, 1998}; ^{Moctezuma 2021}; ^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (^{Pablo-Cea, 2021: 20}; ^{Pablo-Cea, Deloya, MacGregor-Fors, Navarrete-Heredia, & Espinosa-de los Monteros, 2022: 378}); La Libertad: Parque Nacional Walter Thilo Deininger (^{Fuentes, 1998: 29}; ^{Horgan, 2008: 2968}) (Fig. 11).

Years of collection: 1979, 1995-1998, 2018.

Months of collection: September and November.

Topographic zone: coastal mountains, coastal plain.

Elevation range: 135-780 m.

Comments: specimens were captured in baited pitfall traps in Parque Nacional Walter Thilo Deininger (beef carrion, cow and horse dung) (^{Fuentes, 1998}; ^{Horgan, 2008}) and Parque Nacional El Imposible (human dung) (^{Pablo-Cea, 2021}).

Onthophagus (Onthophagus) crinitus Harold, 1869 Distribution: Mexico, Belize, El Salvador, Nicaragua, Costa Rica, and Colombia (^{Pulido-Herrera & Zunino, 2007}; ^{Schoolmeesters, 2022}).

Locality records: no specific department: no specific locality (^{Pulido-Herrera & Zunino, 2007: 97}).

Years of collection: no data.

Months of collection: no data.

Topographic zone: no data.

Elevation range: no data.

Onthophagus (Onthophagus) cyanellus Bates, 1887 Distribution: Mexico, Guatemala, El Salvador, Nicaragua, Honduras, Costa Rica, Panama, Colombia, and Ecuador (^{Pulido-Herrera & Zunino, 2007}; ^{Schoolmeesters, 2022}).

Locality records: no specific department: no specific locality (^{Pulido-Herrera & Zunino, 2007: 102}).

Years of collection: no data.

Months of collection: no data.

Topographic zone: no data.

Elevation range: no data.

Onthophagus (Onthophagus) hoepfneri Harold, 1869 Distribution: USA, Mexico, El Salvador, Nicaragua, and Costa Rica (^{Fuentes, 1998}; ^{Schoolmeesters, 2022}). Locality records: La Libertad: Parque Nacional Walter Thilo Deininger (^{Fuentes, 1998: 29}); La Paz: Comalapa (La Providencia) (^{Horgan, 2001: 106}, ^{2002: 30}, ^{2008: 2969}) (Fig. 11).

Years of collection: 1995-1998.

Months of collection: May and August.

Topographic zone: coastal plain.

Elevation range: 40-135 m.

Comments: specimens were captured in baited pitfall traps in Parque Nacional Walter Thilo Deininger (cow and horse dung) (^{Fuentes, 1998}) and Comalapa (beef carrion, cow and horse dung) (^{Horgan, 2001}).

Onthophagus (Onthophagus) incensus Say, 1835 Distribution: USA, Mexico, Guatemala, El Salvador, Costa Rica, Panama, Venezuela, Ecuador, Uruguay, and Dominican Republic (^{Schoolmeesters, 2022}).

Locality records: Santa Ana: Parque Nacional Montecristo [probably Los Planes] (^{Howden & Cartwright, 1963: 26}); Chalatenango: La Montañona (MUHNES); Morazán: Cerro Perquín (^{Horgan, 2001: 106}) (Fig. 11).

Years of collection: 1996, 1997.

Months of collection: May and August.

Topographic zone: northern mountains.

Elevation range: 1,205-1,890 m.

Comments: Berry and ^{Salazar-Vaquero (1957)} reported the species in San Salvador: Ilopango and Cuscatlán: El Rosario, but they probably misidentified the specimens of an undescribed species later named O. batesi. Onthophagus incensus is found at elevations above 800 m (^{Kohlmann & Solís, 2001}). Specimens were captured in pitfall traps baited with cow and horse dung in Cerro Perquín (^{Horgan, 2001}). Onthophagus (Onthophagus) landolti Harold, 1880

Distribution: Mexico, Guatemala, El Salvador, Nicaragua, Costa Rica, Panama, Colombia, and Venezuela (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El

Imposible (San Benito) (^{Pablo-Cea, 2014: 46}; ^{Pablo-Cea et al., 2020: 6}) (San Francisco Menéndez) (MUHNES, RDCC); Santa Ana: Parque Nacional Montecristo (San José Ingenio) (MUHNES, RDCC); La Libertad: Los Chorros (RDCC); Parque Nacional Walter Thilo Deininger (^{Fuentes, 1998: 29}; ^{Horgan, 2008: 2968}); La Paz: Comalapa (La Providencia) (^{Fuentes, 1998: 29}; ^{Horgan, 2002: 30}) (Fig. 11); no specific department: no specific locality (^{Howden & Young, 1981: 113}).

Years of collection: 1979, 1999, 2013.

Months of collection: May-November.

Topographic zone: northern mountains, coastal mountains, coastal plain.

Elevation range: 40-810 m.

Comments: specimens were captured in baited pitfall traps in Parque Nacional Walter Thilo Deininger (beef carrion, cow and horse dung) (^{Horgan, 2008}), Parque Nacional El Imposible (chicken carrion, human dung) (^{Pablo-Cea, 2014}), and Comalapa (cow and horse dung) (^{Fuentes, 1998}; ^{Horgan, 2002}).

Onthophagus (Onthophagus) marginicollis Harold, 1880 Distribution: Mexico, Guatemala, El Salvador, Nicaragua, Costa Rica, Panama, Colombia, Venezuela,

Guyana, Peru, and Cuba (^{Schoolmeesters, 2022}). Locality records: La Libertad: Parque El Bicentenario (^{Pablo-Cea, 2021: 20}; ^{Pablo-Cea, Deloya, MacGregor-Fors, & Navarrete-Heredia, 2022: 277}; ^{Pablo-Cea, Deloya, MacGregor-Fors, Navarrete-Heredia et al., 2022: 378}); Parque Nacional Walter Thilo Deininger (^{Horgan, 2008: 2968}); La Paz: Comalapa (MUHNES); (La Providencia) (^{Fuentes, 1998: 29}; ^{Horgan, 2001: 106}, ^{2002: 30}, ^{2008: 2969}); Morazán: Jocoaitique (El Rodeo) (^{Fuentes, 1998: 29}; ^{Horgan, 2001: 106}) (Fig. 12); no specific department: no specific locality (^{Howden & Young, 1981: 108}).

Years of collection: 1995-1998, 2019.

Months of collection: January-May, and August.

Topographic zone: northern mountains, coastal mountains, coastal plain.

Elevation range: 40-870 m.

Comments: specimens were captured in baited pitfall traps in Comalapa (beef carrion, cow and horse dung) (^{Horgan, 2001}, ²⁰⁰⁸), Jocoaitique (cow and horse dung) (^{Fuentes, 1998}), Parque Nacional Walter Thilo Deininger (cow dung) (^{Horgan, 2008}), and Parque El Bicentenario (human dung) (^{Pablo-Cea, 2021}).

Onthophagus (Onthophagus) salvadorensis^{Zunino & Halffter, 1988}

Distribution: El Salvador (^{Schoolmeesters, 2022}).

Locality records: Santa Ana: Parque Nacional Montecristo (NE of Metapán) (Bosque Nebuloso) (^{Zunino & Halffter, 1988: 120}; ^{Pulido-Herrera & Zunino, 2007: 115}) (Fig. 12).

Years of collection: 1971.

Months of collection: May.

Topographic zone: northern mountains.

Elevation range: 2,360 m.

Comments: the species is precinctive to El Salvador.

Tribe Phanaeini Hope, 1838 Coprophanaeus Olsoufieff, 1924

Coprophanaeus (Coprophanaeus) boucardi (Nevinson, 1891)

Distribution: El Salvador, Honduras, Nicaragua, and Costa Rica (^{Schoolmeesters, 2022}).

Locality records: La Libertad: Parque El Bicentenario (^{Pablo-Cea, 2021: 20}; ^{Pablo-Cea, Deloya, MacGregor-Fors, & Navarrete-Heredia, 2022: 277}; ^{Pablo-Cea, Deloya, MacGregor-Fors, Navarrete-Heredia et al., 2022: 378}); Santa Tecla (^{Edmonds & Zidek, 2010: 68}); San Salvador: Lago de Ilopango (^{Edmonds & Zidek, 2010: 68}); San Salvador (^{Edmonds & Zidek, 2010: 68}) (Fig. 12).

Years of collection: 2018-2019.

Months of collection: March-October.

Topographic zone: interior valley, coastal mountains.

Elevation range: 510-900 m.

Comments: specimens were captured in pitfall traps baited with squid carrion (^{Pablo-Cea, 2021}). Coprophanaeus (Coprophanaeus) corythus (Harold, 1863)

Distribution: Mexico, Guatemala, Belize, El Salvador, Honduras, Nicaragua, Costa Rica, Panama, Colombia, and Venezuela (^{Fuentes, 1998}; ^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (^{Pablo-Cea, 2014: 46}; ^{Pablo-Cea et al., 2020: 6}); La Libertad: Parque El Bicentenario (^{Pablo-Cea, 2021: 20}; ^{Pablo-Cea, Deloya, MacGregor-Fors, & Navarrete-Heredia, 2022: 277}; ^{Pablo-Cea, Deloya, MacGregor-Fors, Navarrete-Heredia et al., 2022: 378}); Parque Nacional Walter Thilo Deininger (^{Horgan, 2008: 2968}); Morazán: Cerro Perquín (^{Fuentes, 1998: 29}; ^{Horgan, 2001: 106}); Jocoaitique (El Rodeo) (^{Horgan, 2001: 106}) (Fig. 12).

Years of collection: 1997, 2013, 2018-2019.

Months of collection: April-November.

Topographic zone: northern mountains, coastal mountains, coastal plain.

Elevation range: 135-1,205 m.

Comments: reported as Coprophanaeus telamon (Erichson) by ^{Fuentes (1998)}, as C. telemon [sic] by ^{Horgan (2001)}, and as C. telamon telamon by ^{Horgan (2008)}. Specimens were captured in baited pitfall traps in Parque Nacional Walter Thilo Deininger (cow dung, beef carrion, and fruit) (^{Horgan, 2008}), Parque Nacional El Imposible (carrion chicken, human dung) (^{Pablo-Cea, 2014}), Parque El Bicentenario (squid carrion, human dung) (^{Pablo-Cea, 2021}), and Jocoaitique and Cerro Perquín (cow dung) (^{Horgan, 2001}).

Phanaeus MacLeay, 1819

Phanaeus (Notiophanaeus) pacificus^{Moctezuma & Halffter, 2021}

Distribution: Mexico, Guatemala, and El Salvador (^{Moctezuma & Halffter, 2021}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (^{Pablo-Cea, 2014: 46}; ^{Pablo-Cea et al., 2020: 6}); La Libertad: Parque El Bicentenario (^{Pablo-Cea, 2021: 20}; ^{Pablo-Cea, Deloya, MacGregor-Fors, & Navarrete-Heredia, 2022: 277}; ^{Pablo-Cea, Deloya, MacGregor-Fors, Navarrete-Heredia et al., 2022: 378}); Parque Nacional Walter Thilo Deininger (^{Fuentes, 1998: 29}; ^{Horgan, 2008: 2968}; ^{Moctezuma & Halffter, 2021: 33}); San Andrés (MUHNES); Santa Tecla (^{Edmonds, 1994: 101}); San Salvador: Tonacatepeque (^{Fuentes, 2009: 22}); Cuscatlán: Área Natural Protegida Colima (Cerrón Grande) (J. Pablo, personal observation); Morazán: Cerro Perquín (^{Fuentes, 1998: 29}; ^{Horgan, 2001: 106}); Jocoaitique (El Rodeo) (^{Horgan, 2001: 106}) (Fig. 12).

Years of collection: 1976, 1995-1998, 2012-2013, 2018.

Months of collection: May-November.

Topographic zone: northern mountains, interior valley, coastal mountains, coastal plain.

Elevation range: 135-1,205 m.

Comments: reported as Phanaeus endymiun [sic] in ^{Fuentes (2009)} and as P. endymion Harold by ^{Edmonds (1994)}, ^{Horgan (2001}, ²⁰⁰⁸⁾, ^{Pablo-Cea (2014)}, and ^{Pablo-Cea et al. (2021)}. The specimens in MUHNES are labeled as P. endymion. Specimens were captured in baited pitfall traps in Parque Nacional Walter Thilo Deininger (beef carrion, cow dung, fruit) (^{Horgan, 2008}), Parque Nacional El Imposible (carrion chicken, human dung) (^{Pablo-Cea, 2021}), Comalapa, Jocoaitique, and Cerro Perquín (cow and horse dung) (^{Horgan, 2001}), and Tonacatepeque and Colima (cow dung) (^{Fuentes, 2009}; J. Pablo, personal observation).

Phanaeus (Phanaeus) excelsus Bates, 1889

Distribution: Guatemala, El Salvador, Honduras, Nicaragua, and Costa Rica (^{Schoolmeesters, 2022}). Locality records: Santa Ana: San Diego (^{Edmonds, 1994: 100}); La Libertad: Quezaltepeque (^{Edmonds, 1994: 100}); San Salvador: Tonacatepeque (^{Fuentes, 2009: 22}); Cuscatlán: Área Natural Protegida Colima (Cerrón Grande) (J. Pablo, personal observation); La Paz: Comalapa (Aeropuerto) (MUHNES); (La Providencia) (^{Fuentes, 1998: 29} ; ^{Horgan, 2001: 106}, ^{2002: 30}, ^{2008: 2969}); Usulután: Usulután (^{Edmonds, 1994: 100}); La Unión: Playa El Icacal (^{Edmonds, 1994: 100}); Volcán de Conchagua (^{Edmonds, 1994: 100}) (Fig. 12).

Years of collection: 1995-1998.

Months of collection: May and August.

Topographic zone: interior valley, coastal mountains, coastal plain.

Elevation range: 15-1,190 m.

Comments: reported as Phanaeus demon Laporte in all references, but this species only occurs in Mexico and Guatemala (^{Edmonds & Zidek 2012}; V. Moctezuma, personal communication). Specimens were captured in baited pitfall traps in Comalapa (beef carrion, cow and horse dung) (^{Horgan, 2001}, ²⁰⁰⁸) and Tonacatepeque, Colima (cow dung) (^{Fuentes, 2009}; J. Pablo, personal observation).

Phanaeus (Phanaeus) eximius Bates, 1887

Distribution: Guatemala, El Salvador, Honduras, Nicaragua, and Costa Rica (^{Schoolmeesters, 2022}). Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (^{Pablo-Cea, 2014: 46}; ^{Pablo-Cea et al., 2020: 6}); La Libertad: Parque Nacional Walter Thilo Deininger (^{Fuentes, 1998: 29}; ^{Horgan, 2008: 2968}); Morazán: Cerro Perquín (^{Horgan, 2001: 106}); Jocoaitique (El Rodeo) (^{Fuentes, 1998: 29}; ^{Horgan, 2001: 106}); no specific department: no specific locality (^{Berry, 1959b: 61}) (Fig. 12).

Years of collection: 1995-1998, 2013.

Months of collection: May, June, and August.

Topographic zone: northern mountains, coastal mountains, coastal plain.

Elevation range: 135-1,205 m.

Comments: specimens were captured in baited pitfall traps in Parque Nacional Walter Thilo Deininger (beef carrion, cow dung, fruit) (^{Horgan, 2008}), Cerro Perquín (cow and horse dung) (^{Horgan, 2001}), Parque Nacional El Imposible (human dung) (^{Pablo-Cea, 2014}), and Colima and Jocoaitique (cow dung) (^{Horgan, 2001}; ^{Fuentes, 2009}; J. Pablo, personal observation).

Phanaeus (Phanaeus) wagneri Harold, 1863 Distribution: Mexico, Guatemala, El Salvador,

Honduras, Nicaragua, and Costa Rica (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (^{Pablo-Cea, 2014: 46}; ^{Pablo-Cea et al., 2020: 6}); Chalatenango: La Palma (^{Edmonds, 1994: 105}); La Libertad: Parque Nacional Walter Thilo

Deininger (^{Fuentes, 1998: 29}; ^{Horgan, 2008: 2968}); Quezaltepeque (^{Edmonds, 1994: 105}); San Salvador: San Salvador (^{Edmonds, 1994: 105}) (Fig. 12).

Years of collection: 1976-1979, 1995-1997, 2013.

Months of collection: May, June, August, and October.

Topographic zone: northern mountains, interior valley, coastal mountains, coastal plain.

Elevation range: 135-1,005 m.

Comments: specimens were captured in baited pitfall traps in Parque Nacional Walter Thilo Deininger (cow dung) (^{Fuentes, 1998}) and Parque Nacional El Imposible (human dung) (^{Pablo-Cea, 2014}).

Tribe Sisyphini Mulsant, 1842

Sisyphus Latreille, 1807

Sisyphus (Sisyphus) mexicanus Harold, 1863 Distribution: Mexico and El Salvador (^{Hogan 2008};

^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (^{Pablo-Cea, 2014: 46}, ^{2021: 20};

Pablo-Cea et al., 2020: 6); La Libertad: Parque Nacional Walter Thilo Deininger (Horgan, 2008: 2968) (Fig. 12).

Years of collection: 2013, 2019.

Months of collection: May, August, and November.

Topographic zone: coastal mountains, coastal plain.

Elevation range: 135-780 m.

Comments: this species is rare in El Salvador. Only 3 specimens have been collected in the country, all in baited pitfall traps, 1 in Parque Nacional Walter Thilo Deininger (cow dung) (^{Horgan, 2008}) and 2 in Parque Nacional El Imposible (human dung) (^{Pablo-Cea, 2014}, ²⁰²¹).

Subfamily Melolonthinae Leach, 1819 Tribe Diplotaxini Kirby, 1837 Diplotaxis Kirby, 1837 Diplotaxis angustula Moser, 1918

Distribution: Mexico, Guatemala, and El Salvador (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (^{Pablo-Cea, 2021: 21}); La Libertad: Santa Tecla (^{Vaurie, 1960: 415}); San Salvador: San Salvador (^{Vaurie, 1960: 415}) (Fig. 12).

Years of collection: 2018-2019.

Months of collection: January, April-July, and September.

Topographic zone: interior valley, coastal mountains.

Elevation range: 685-900 m.

Comments: collected with UV light traps in Parque Nacional El Imposible.

Diplotaxis brevipilosa Moser, 1918

Distribution: Guatemala (^{Schoolmeesters, 2022}) and El Salvador.

Locality records: Cabañas: Cinquera (RDCC) (Fig. 13).

Years of collection: 2001.

Months of collection: June.

Topographic zone: interior valley.

Elevation range: 385 m.

Comments: new country record.

Diplotaxis cavifrons Moser, 1918

Distribution: Mexico, Guatemala (^{Schoolmeesters, 2022}), and El Salvador.

Locality records: Santa Ana: Parque Nacional Montecristo (Los Planes) (RDCC) (Fig. 13).

Years of collection: 1999.

Months of collection: June.

Topographic zone: northern mountains.

Elevation range: 1,890 m.

Comments: new country record.

Diplotaxis crinigera Bates, 1888

Distribution: Mexico, Guatemala, and El Salvador (^{Schoolmeesters, 2022}).

Locality records: San Salvador: San Salvador (^{Vaurie, 1960: 176}); La Unión: Volcán de Conchagua (^{Vaurie, 1960: 176}) (Fig. 13).

Years of collection: no data.

Months of collection: no data.

Topographic zone: interior valley, coastal mountains.

Elevation range: 685-1,190 m.

Diplotaxis denigrata Bates, 1889

Distribution: Mexico, El Salvador, Honduras, Nicaragua, and Costa Rica (^{Schoolmeesters, 2022}).

Locality records: La Unión: Volcán de Conchagua ^{(Vaurie, 1960: 419}) (Fig. 13).

Months of collection: no data.

Topographic zone: coastal mountains.

Elevation range: 1,190 m. Diplotaxis macrotarsus Vaurie, 1960

Distribution: Mexico, Guatemala, and El Salvador (^{Schoolmeesters, 2022}).

Locality records: San Salvador: San Salvador (^{Vaurie, 1960: 422}); La Unión: Volcán de Conchagua (^{Vaurie, 1960: 422}) (Fig. 13).

Years of collection: 1957, 1958.

Months of collection: May and June.

Topographic zone: interior valley, coastal mountains.

Elevation range: 685-1,190 m.

Comments: some specimens were found in debris outside a leafcutter ant nest (^{Vaurie, 1960}).

Diplotaxis mistura Vaurie, 1960

Distribution: Mexico, El Salvador, and Honduras (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (^{Pablo-Cea, 2021: 21}; ^{Pablo-Cea, Deloya, MacGregor-Fors, Navarrete-Heredia, & Espinosa-de los Monteros, 2022: 378}); Santa Ana: Metapán (^{Vaurie, 1960: 401}); Parque Nacional Montecristo (La Torre, Majaditas) (^{Serrano-Chicas, 2019: 39, 40}); San Salvador: San Salvador (^{Vaurie, 1960: 401}); Cabañas: Cinquera (RDCC); La Unión: Volcán de Conchagua (^{Vaurie, 1960: 423}) (Fig. 13).

Years of collection: 1957-1958, 2000, 2018-2019.

Months of collection: January, April-July, and September.

Topographic zone: northern mountains, interior valley, coastal mountains.

Elevation range: 385-1,450 m.

Comments: collected with UV light traps in Parque Nacional El Imposible (^{Pablo-Cea, 2021}). The species was described from specimens collected at lights in San Salvador (^{Vaurie, 1960}).

Diplotaxis ohausi Moser, 1921

Distribution: Mexico, Guatemala, El Salvador, and Honduras (^{Vaurie, 1960}; ^{Schoolmeesters, 2022}).

Locality records: San Salvador: San Salvador (^{Vaurie, 1960: 177}) (Fig. 13).

Years of collection: 1957-1958.

Months of collection: May.

Topographic zone: interior valley.

Elevation range: 685 m. Diplotaxis pacata LeConte, 1856

Distribution: USA, Mexico, and El Salvador (^{Schoolmeesters, 2022}).

Locality records: no specific department: no specific locality (^{Vaurie, 1960: 176}).

Years of collection: no data.

Months of collection: no data.

Topographic zone: no data.

Elevation range: no data.

Diplotaxis poropyge Bates, 1887

Distribution: Mexico, Guatemala, El Salvador, Nicaragua, Costa Rica, and Panama (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (EAPZ, RDCC); Chalatenango: Las Ceibas [probably Las Vueltas] (^{Vaurie, 1958: 392}); Cuscatlán: El Rosario (^{Vaurie, 1958: 392}) (Fig. 13).

Years of collection: 1955, 1999-2000.

Months of collection: May.

Topographic zone: interior valley, coastal mountains.

Elevation range: 710-780 m.

Comments:^{Vaurie (1958)} examined 1 female collected at Servicios Técnicos Cafetalera, a locality unknown to us. Diplotaxis puncticollis Moser, 1918

Distribution: Mexico, Guatemala, and El Salvador (^{Schoolmeesters, 2022}).

Locality records: San Salvador: San Salvador (^{Vaurie, 1960: 177}) (Fig. 13).

Years of collection: 1957-1958.

Months of collection: May.

Topographic zone: interior valley.

Elevation range: 685 m. Diplotaxis trapezifera Bates, 1887

Distribution: Mexico, Guatemala, Belize, El Salvador, Honduras, Nicaragua, and Costa Rica (^{Vaurie, 1960}; ^{Schoolmeesters, 2022}).

Locality records: San Salvador: San Salvador (^{Vaurie, 1960: 427}); Cuscatlán: El Rosario (^{Vaurie, 1960: 427}) (Fig. 13).

Years of collection: no data.

Months of collection: no data.

Topographic zone: interior valley.

Elevation range: 685-720 m. Diplotaxis yucateca Vaurie, 1960

Distribution: Mexico, El Salvador, Honduras, and Costa Rica (^{Vaurie, 1960}; ^{Schoolmeesters, 2022}).

Locality records: San Salvador: San Salvador (^{Vaurie, 1960: 428}); La Unión: Volcán de Conchagua (^{Vaurie, 1960: 428}) (Fig. 14).

Years of collection: 1958.

Months of collection: no data.

Topographic zone: interior valley, coastal mountains.

Elevation range: 685-1,190 m.

Tribe Hopliini Latreille, 1829

Hoplia Illiger, 1803

Hoplia (Hoplia) guatemalensis Bates, 1887

Distribution: Mexico, Guatemala, and El Salvador (^{Berry, 1959b}; ^{Schoolmeesters, 2022}).

Locality records: no specific department: no specific locality (^{Berry, 1959b: 40}).

Years of collection: no data.

Months of collection: no data.

Topographic zone: no data.

Elevation range: no data.

Tribe Macrodactylini Kirby, 1837 Isonychus Mannerheim, 1828 Isonychus ocellatus Burmeister, 1855

Distribution: Mexico, Guatemala, El Salvador, and Costa Rica (^{Serrano-Chicas, 2019}; ^{Schoolmeesters, 2022}).

Locality records: Santa Ana: Parque Nacional Montecristo (La Torre, Majaditas) (^{Serrano-Chicas, 2019: 39, 40}) (Fig. 14).

Years of collection: 2019.

Months of collection: April.

Topographic zone: northern mountains.

Elevation range: 1,070-1,450 m.

Isonychus pictus Sharp, 1877

Distribution: Nicaragua, Costa Rica, Panama (^{Schoolmeesters, 2022}), and El Salvador.

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (RDCC) (Fig. 14).

Years of collection: 2000.

Months of collection: May.

Topographic zone: coastal mountains.

Elevation range: 780 m.

Comments: new country record.

Macrodactylus Dejean, 1821 Macrodactylus costulatus Bates, 1887

Distribution: Mexico, Guatemala, Belize, El Salvador, Honduras, Nicaragua, Costa Rica, and Panama (^{Schoolmeesters, 2022}).

Locality records: Chalatenango: La Palma (^{Arce-Pérez, 2008: 55}; ^{Arce-Pérez & Morón, 2020: 572}) (Fig. 14).

Years of collection: 1955.

Months of collection: June.

Topographic zone: northern mountains.

Elevation range: 1,005 m.

Macrodactylus hondurensis^{Arce-Pérez & Morón, 2005} Distribution: El Salvador and Honduras (^{Arce-Pérez & Morón, 2020}; ^{Schoolmeesters, 2022}).

Locality records: Santa Ana: Parque Nacional Montecristo (Bosque Nebuloso) (^{Arce-Pérez & Morón, 2020: 575}); Chalatenango: Cerro El Pital (EAPZ) (Fig. 14).

Years of collection: 1971, 1994.

Months of collection: May and July.

Topographic zone: northern mountains.

Elevation range: 2,360-2,660 m.

Macrodactylus lineatus Chevrolat, 1834

Distribution: Mexico and El Salvador (^{Berry, 1959b}; ^{Schoolmeesters, 2022}).

Locality records: no specific department: no specific locality (^{Berry, 1959b: 47}).

Years of collection: no data.

Months of collection: no data.

Topographic zone: no data.

Elevation range: no data.

Macrodactylus montanus^{Arce-Pérez & Morón 2000}Distribution: Mexico, El Salvador, Honduras, and Nicaragua (^{Schoolmeesters, 2022}).

Locality records: Santa Ana: Parque Nacional Cerro Verde (^{Arce-Pérez, 2008: 94}); Parque Nacional Montecristo (Los Planes) (^{Serrano-Chicas, 2019: 39, 40}) (Fig. 14).

Years of collection: 1971, 2019.

Months of collection: May.

Topographic zone: northern mountains, coastal mountains.

Elevation range: 1,890-2,020 m.

Macrodactylus sericeicollis Bates, 1887

Distribution: Mexico, Guatemala, El Salvador, Honduras, and Nicaragua (^{Schoolmeesters, 2022}).

Locality records: Santa Ana: Parque Nacional Cerro Verde (^{Arce-Pérez, 2008: 115}) (Fig. 14).

Years of collection: 1963.

Months of collection: June.

Topographic zone: coastal mountains.

Elevation range: 2,020 m.

Macrodactylus zunilensis Bates, 1887

Distribution: Mexico, Guatemala, and El Salvador (^{Arce-Pérez & Morón, 2020}; ^{Schoolmeesters, 2022}).

Locality records: no specific department: no specific locality (^{Arce-Pérez, 2008: 153}).

Years of collection: no data.

Months of collection: no data.

Topographic zone: no data.

Elevation range: no data.

Tribe Rhizotrogini Burmeister, 1855 Chlaenobia Blanchard, 1850 Chlaenobia aequata Bates, 1887

Distribution: USA, Mexico, Guatemala, El Salvador, Honduras, Nicaragua, and Costa Rica (^{Evans & Smith, 2009}; ^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (EAPZ, RDCC); San Salvador: San Salvador (^{Morón, 2006: 37}) (Fig. 14); no specific department: no specific locality (^{Morón, 1994: 71}).

Years of collection: 1958, 1999-2000.

Months of collection: May.

Topographic zone: interior valley, coastal mountains.

Elevation range: 685-780 m.

Comments: reported as Phyllophaga (Chlaenobia) aequata by ^{Morón (2006)}.

Chlaenobia chiapensis Chapin, 1935

Distribution: Mexico, Guatemala, El Salvador, Honduras, Nicaragua, and Costa Rica (^{Schoolmeesters, 2022}).

Locality records: San Salvador: San Salvador (^{Morón, 2006: 37}) (Fig. 14).

Years of collection: no data.

Months of collection: no data.

Topographic zone: interior valley.

Elevation range: 685 m.

Comments: reported as Phyllophaga (Chlaenobia) chiapensis by ^{Morón (2006)}.

Chlaenobia instabilis (Blackwelder, 1944)

Distribution: Mexico, Guatemala, and El Salvador (^{Rivera-Gasperín & Morón, 2017}).

Locality records: no specific department: no specific locality (^{Rivera-Gasperín & Morón, 2017: 606}).

Years of collection: no data.

Months of collection: no data.

Topographic zone: no data.

Elevation range: no data. Chlaenobia latipes Bates, 1888

Distribution: Mexico, Guatemala, and El Salvador (^{Evans, 2003}; ^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Turín (^{Mendoza, 1994: 16}); Sonsonate: Izalco (^{Mendoza, 1994: 16}); Chalatenango: La Palma (^{Morón, 2006: 30}); San Salvador: San Salvador (^{Morón, 2006: 37}) (Fig. 14); no specific department: no specific locality (^{Andrews et al., 1979: 7}).

Years of collection: no data.

Months of collection: June and July.

Topographic zone: northern mountains, interior valley, coastal mountains.

Elevation range: 470-1,005 m.

Comments: reported as Phyllophaga (Chlaenobia) latipes by ^{Morón (2006)}. The species is considered a part of the “gallina ciega” complex in cultivated corn in El Salvador, and it is associated with cultivated pineapple (^{Mendoza, 1994}).

Chlaenobia scabripyga Bates, 1888

Distribution: Mexico, Guatemala, El Salvador, and Costa Rica (^{Rivera-Gasperín & Morón, 2017}).

Locality records: no specific department: no specific locality (^{Evans & Smith, 2009: 128}).

Years of collection: no data.

Months of collection: no data.

Topographic zone: no data.

Elevation range: no data.

Comments: reported as Phyllophaga (Chlaenobia) scabripyga by ^{Evans & Smith (2009)}.

Chlaenobia solanophaga (^{Morón, 1988}) Distribution: El Salvador (^{Morón 1988}).

Locality records: Santa Ana: Parque Nacional Montecristo (Majaditas, San José Ingenio) (^{Serrano-Chicas, 2019: 39, 40}); Chalatenango: Las Pilas (^{Morón, 1988: 60}) (Fig. 15).

Years of collection: 1978-1979, 1984, 1999, 2019.

Months of collection: February, March, April, May (adults), and July (immatures).

Topographic zone: northern mountains.

Elevation range: 810-1,700 m.

Comments: described as Phyllophaga solanophaga by ^{Morón (1988)}. Reported as Phyllophaga zunilensis (Bates) by ^{King (1994)}, but ^{Morón (1988)} pointed out this species does not occur in El Salvador. The larvae are pests in cultivated potato (^{Morón, 1988}; ^{King, 1994}). Chlaenobia solanophaga is precinctive to El Salvador.

Chlaenobia tumulosa Bates, 1888

Distribution: Mexico, Belize, Guatemala, El Salvador, Honduras, Nicaragua, and Costa Rica (^{Saunders et al., 1998}; ^{Evans & Smith, 2009}; ^{Rivera-Gasperín & Morón, 2017}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (^{Pablo-Cea, 2021: 21}; ^{Pablo-Cea, Deloya, MacGregor-Fors, Navarrete-Heredia, & Espinosa-de los Monteros, 2022: 378}); Santa Ana: Parque Nacional Montecristo (Los Planes) (RDCC); (Majaditas, San José Ingenio) (^{Serrano-Chicas, 2019: 40, 41}); La Libertad: Parque El Bicentenario (^{Pablo-Cea, 2021: 21}; ^{Pablo-Cea, Deloya, MacGregor-Fors, & Navarrete-Heredia, 2022: 277}; ^{Pablo-Cea, Deloya, MacGregor-Fors, Navarrete-Heredia et al., 2022: 378}); no specific department: no specific locality (^{Morón, 1994: 73}) (Fig. 15).

Years of collection: 1999-2000, 2019.

Months of collection: March-July.

Topographic zone: northern mountains, coastal mountains.

Elevation range: 780-1,890 m.

Comments: reported as Phyllophaga (Chlaenobia) tumulosa by ^{King (1994)}, ^{Morón (1994)}, ^{Evans & Smith (2009)}, and ^{Serrano-Chicas (2019)}.

Chlaenobia vexata (Horn, 1885)

Distribution: USA, Mexico, Guatemala (^{Rivera-Gasperín & Morón, 2017}), and El Salvador.

Locality records: Santa Ana: Parque Nacional Montecristo (Los Planes) (RDCC) (Fig. 15).

Years of collection: 1999.

Months of collection: May.

Topographic zone: northern mountains.

Elevation range: 1,890 m.

Comments: new country record.

Phyllophaga Harris, 1826

Phyllophaga (Phyllophaga) baneta Saylor, 1943 Distribution: Guatemala, Honduras, Nicaragua

(^{Schoolmeesters, 2022}), and El Salvador.

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (RDCC) (Fig. 15).

Years of collection: 1999.

Months of collection: May.

Topographic zone: coastal mountains.

Elevation range: 780 m.

Comments: new country record. Phyllophaga (Phyllophaga) caraga Saylor, 1943

Distribution: El Salvador, Nicaragua, Costa Rica, and Panama (^{Serrano-Chicas, 2019}; ^{Schoolmeesters, 2022}).

Locality records: Santa Ana: Parque Nacional Montecristo (Majaditas) (^{Serrano-Chicas, 2019: 39, 40}) (Fig. 15).

Years of collection: 2019.

Months of collection: March.

Topographic zone: northern mountains.

Elevation range: 1,070 m.

Phyllophaga (Phyllophaga) colimana (Moser, 1921) Distribution: Mexico (^{Schoolmeesters, 2022}) and El Salvador.

Locality records: La Libertad: Volcán de San Salvador (El Boquerón) (EAPZ) (Fig. 15).

Years of collection: 1971.

Months of collection: May.

Topographic zone: coastal mountains.

Elevation range: 1,845 m.

Comments: new country record.

Phyllophaga (Phyllophaga) dasypoda (Bates, 1888) Distribution: Mexico, Guatemala, El Salvador, and Costa Rica (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Atiquizaya (^{Mendoza, 1994: 16}); Parque Nacional El Imposible (San Benito) (RDCC); San Salvador: San Salvador (EAPZ) (Fig. 15).

Years of collection: 1978, 1999-2000.

Months of collection: February, and May-July.

Topographic zone: interior valley, coastal mountains.

Elevation range: 590-780 m.

Comments: the species is considered a part of the “gallina ciega” complex in cultivated corn in El Salvador (^{Mendoza, 1994}).

Phyllophaga (Phyllophaga) elenans Saylor, 1938 Distribution: Belize, Guatemala, El Salvador, Honduras, Nicaragua, and Costa Rica (^{Schoolmeesters, 2022}). Locality records: Ahuachapán: Atiquizaya (^{Mendoza, 1994: 16}) (Fig. 15).

Years of collection: no data.

Months of collection: no data.

Topographic zone: interior valley.

Elevation range: 590 m.

Comments: the species is considered a part of the “gallina ciega” complex in cultivated corn (^{Mendoza, 1994}).

Phyllophaga (Phyllophaga) fulviventris (Moser, 1918) Distribution: Mexico and El Salvador (King, 1994; Schoolmeesters, 2022).

Locality records: no specific department: no specific locality (^{King, 1994: 37}; ^{Morón, 1994: 71}).

Years of collection: no data.

Months of collection: no data.

Topographic zone: no data.

Elevation range: no data.

Phyllophaga (Phyllophaga) menetriesii (Blanchard, 1851) Distribution: Mexico, Guatemala, El Salvador, Honduras, Nicaragua, Costa Rica, Panama, Colombia, and Venezuela (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Atiquizaya (^{Mendoza, 1994: 16}); Parque Nacional El Imposible (San Benito) (EAPZ, RDCC); Santa Ana: Parque Nacional Montecristo (Majaditas, San José Ingenio) (^{Serrano-Chicas, 2019: 39, 40}); Sonsonate: Izalco (^{Mendoza, 1994: 16}); no specific department: no specific locality (^{Berry, 1959b: 62}) (Fig. 15).

Years of collection: 1999, 2019.

Months of collection: March-May.

Topographic zone: northern mountains, interior valley, coastal mountains, coastal plain.

Elevation range: 470-1,070 m.

Comments: the species is considered a part of the “gallina ciega” complex in cultivated corn in El Salvador, and it is associated with cultivated pineapple (^{Mendoza, 1994}).

Phyllophaga (Phyllophaga) parvisetis (Bates, 1888) Distribution: Mexico, Belize, Guatemala, El Salvador, Honduras, and Nicaragua (^{Schoolmeesters, 2022}). Locality records: no specific department: no specific locality (^{Morón, 1994: 72}; ^{Saunders et al., 1998: 136}).

Years of collection: no data.

Months of collection: no data.

Topographic zone: no data.

Elevation range: no data.

Phyllophaga (Phyllophaga) setidorsis (Bates, 1888) Distribution: Mexico, El Salvador, and Costa Rica

(^{Evans, 2003}; ^{Serrano-Chicas, 2019}).

Locality records: Santa Ana: Parque Nacional Montecristo (Majaditas) (^{Serrano-Chicas, 2019: 39, 40}) (Fig. 15).

Years of collection: 2019.

Months of collection: March.

Topographic zone: northern mountains.

Elevation range: 1,070 m.

Phyllophaga (Phyllophaga) valeriana Saylor, 1934 Distribution: Guatemala, El Salvador, Nicaragua, and Costa Rica (^{Morón, 1994}; ^{Schoolmeesters, 2022}). Locality records: Santa Ana: Parque Nacional Montecristo (Majaditas) (Serrano-Chicas, 2019: 39-40) (Fig. 16); no specific department: no specific locality (^{Morón, 1994: 72}).

Years of collection: 2019.

Months of collection: March and April.

Topographic zone: northern mountains.

Elevation range: 1,070 m.

Comments: reported as Phyllophaga sanjosicola by ^{Saunders et al. (1998)}, ^{Evans (2003)}, and ^{Serrano-Chicas (2019)}.

Phyllophaga (Phyllophaga) vicina (Moser, 1918) Distribution: Guatemala, El Salvador, Nicaragua,

Costa Rica, and Panama (^{Mendoza, 1994}; ^{Schoolmeesters, 2022})

Locality records: Ahuachapán: Atiquizaya (Izcaquillo)

(^{Mendoza, 1994: 16}); Turín (^{Mendoza, 1994: 16}); San Salvador: San Salvador (^{Morón, 2006: 37}) (Fig. 16).

Years of collection: 1958, 2019.

Months of collection: April-July.

Topographic zone: interior valley.

Elevation range: 630-700 m.

Phyllophaga (Phyllophaga) yucateca (Bates, 1889) Distribution: Mexico, Guatemala, Belize, El Salvador, Honduras, Nicaragua, and Costa Rica (^{Morón, 1994}; ^{Evans, 2003})

Locality records: no specific department: no specific

locality (^{Morón, 1994: 73}).

Years of collection: no data.

Months of collection: no data.

Topographic zone: no data.

Elevation range: no data.

Phyllophaga (Phytalus) cometes (Bates, 1887) Distribution: Mexico, Guatemala, El Salvador,

Nicaragua, Costa Rica, and Panama (^{Serrano-Chicas, 2019}; ^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (RDCC); Santa Ana: Parque Nacional Montecristo (Majaditas, San José Ingenio) (^{Serrano-Chicas, 2019: 39, 40}) (Fig. 16).

Years of collection: 1999, 2019.

Months of collection: March-May.

Topographic zone: northern mountains, coastal mountains.

Elevation range: 780-1,070 m. Phyllophaga (Phytalus) guatemala Saylor, 1940

Distribution: Mexico, Guatemala, El Salvador, and Honduras (^{Schoolmeesters, 2022}).

Locality records: Santa Ana: Parque Nacional Montecristo (Bosque Nebuloso) (^{Morón, 2006: 37}) (Fig. 16).

Years of collection: 1971.

Months of collection: May.

Topographic zone: northern mountains.

Elevation range: 2,360 m.

Phyllophaga (Phytalus) obsoleta Blanchard, 1851 Distribution: USA, Mexico, Belize, Guatemala, El Salvador, Honduras, Nicaragua, Costa Rica, Panama, Colombia, and Venezuela (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (^{Pablo-Cea, 2021: 21}; ^{Pablo-Cea, Deloya, MacGregor-Fors, Navarrete-Heredia, & Espinosa-de los Monteros, 2022: 378}); Santa Ana: Metapán (^{Morón, 2006: 30}); Parque Nacional Montecristo (Bosque Nebuloso) (^{Morón, 2006: 30, 37}); (La Torre, Los Planes, Majaditas) (^{Serrano-Chicas, 2019: 39, 40}); Chalatenango: La Palma (^{Morón, 2006: 30, 37}); La Libertad: Parque El Bicentenario (^{Pablo-Cea, 2021: 21; Pablo-Cea, Deloya, MacGregor-Fors, & Navarrete-Heredia, 2022: 277}; ^{Pablo-Cea, Deloya, MacGregor-Fors, Navarrete-Heredia et al., 2022: 378}); Santa Tecla (^{Morón, 2006: 30}); Volcán de San Salvador (El Boquerón) (^{Morón, 2006: 30}); San Salvador: San Salvador (^{Morón, 2006: 29-30, 37}); no specific department: no specific locality (^{Morón, 1994: 72}) (Fig. 16).

Years of collection: 1958-1959, 1971, 1999-2000, 2018-2019.

Months of collection: March-May.

Topographic zone: northern mountains, interior valley, coastal mountains.

Elevation range: 530-2,360 m.

Phyllophaga (Phytalus) pruinosa (Blanchard, 1851) Distribution: México, El Salvador, Nicaragua, and Costa Rica (^{Morón, 2018}; ^{Schoolmeesters, 2022}; ^{Pablo-Cea, 2021}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (^{Pablo-Cea, 2021: 21}; ^{Pablo-Cea, Deloya, MacGregor-Fors, Navarrete-Heredia, & Espinosa-de los Monteros, 2022: 378}) (Fig. 16).

Years of collection: 2018-2019.

Months of collection: April-September.

Topographic zone: coastal mountains.

Elevation range: 780 m.

Phyllophaga (Phytalus) punctuliceps (Bates, 1888) Distribution: Mexico, Guatemala, (^{Schoolmeesters, 2022}), and El Salvador.

Locality records: Santa Ana: Parque Nacional Montecristo (Los Planes) (RDCC) (Fig. 16).

Years of collection: 1999.

Months of collection: May.

Topographic zone: northern mountains.

Elevation range: 1,890 m.

Comments: new country record.

Subfamily Rutelinae MacLeay, 1819 Tribe Anomalini Streubel, 1839

Callistethus Blanchard, 1851 Callistethus granulipygus (Bates, 1887)

Distribution: Guatemala, El Salvador, Nicaragua, Costa Rica, and Panama (^{Berry, 1959b}; ^{Schoolmeesters, 2022}).

Locality records: Santa Ana: Parque Nacional Montecristo (Los Planes) (RDCC) (Fig. 16); no specific department: no specific locality (^{Berry, 1959b: 6}).

Years of collection: 1999.

Months of collection: May.

Topographic zone: northern mountains.

Elevation range: 1,890 m.

Comments: the species was reported as Anomala granulipyga by ^{Berry (1959b)}.

Callistethus multiplicatus Filippini, Galante, & Micó, 2015 Distribution: El Salvador and Costa Rica

(^{Schoolmeesters, 2022}; ^{Pablo-Cea, 2021}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (^{Pablo-Cea, 2021: 21}; ^{Pablo-Cea, Deloya, MacGregor-Fors, Navarrete-Heredia, & Espinosa-de los Monteros, 2022: 378}) (Fig. 16).

Years of collection: 2018-2019.

Months of collection: May-December.

Topographic zone: coastal mountains.

Elevation range: 780 m.

Epectinaspis Blanchard, 1850

Epectinaspis moreletiana (Blanchard, 1850)

Distribution: Mexico, Guatemala, El Salvador, Nicaragua, Panama, and Colombia (^{Schoolmeesters, 2022}).

Locality records: Sonsonate: Izalco (Los Guates Farm) (^{Paucar-Cabrera, 2003: 42}); La Libertad: La Libertad (^{Paucar-Cabrera, 2003: 42}); Santa Tecla (^{Berry & Salazar-Vaquero, 1957: 44}; ^{Paucar-Cabrera, 2003: 42}); San Salvador: Cerro San Jacinto (^{Paucar-Cabrera, 2003: 42}); San Salvador (^{Paucar-Cabrera, 2003: 42}) (Fig. 16).

Years of collection: 1951, 1974.

Months of collection: June and November.

Topographic zone: interior valley, coastal mountains, coastal plain.

Elevation range: 70-980 m.

Comments: reported as Epectinaspis quadripennis Casey by Berry and ^{Salazar-Vaquero (1957)}.

Moroniella Ramírez-Ponce, 2015

Moroniella nitidula nitidula (Blanchard, 1850) Distribution: Guatemala, El Salvador, Honduras, Nicaragua, and Costa Rica (^{Schoolmeesters, 2022}). Locality records: Santa Ana: Hacienda Las Brumas (^{Franz, 1955: 206}); Parque Nacional Cerro Verde (^{Franz, 1955: 206}); Parque Nacional Montecristo (Los Planes) (^{Franz, 1955: 206}; ^{Serrano-Chicas, 2019: 42}); Volcán de Santa Ana (^{Franz, 1955: 206}); Chalatenango: Cerro El Pital (EAPZ); Near Lempa Bridge (Road to La Palma) (^{Franz, 1955: 206}) (Fig. 17).

Years of collection: 1951-1952, 1979, 1994, 2018.

Months of collection: August, October, and November.

Topographic zone: northern mountains, interior valley, coastal mountains.

Elevation range: 245-2,660 m.

Comments: reported as Anomala nitidula by ^{Franz (1955)}.

Strigoderma Burmeister, 1844 Strigoderma castor (Newman, 1838)

Distribution: Mexico, Guatemala, El Salvador, Nicaragua, and Costa Rica (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (RDCC); La Libertad: Santa Tecla (^{Berry & Salazar-Vaquero, 1957: 47}) (Fig. 17).

Years of collection: 1979.

Months of collection: June.

Topographic zone: coastal mountains.

Elevation range: 780-900 m.

Comments: map 3 in Bader (1992, p. 328) shows 1 point that does not correlate with either of the 2 Salvadoran locality records known for the species.

Strigoderma mexicana Blanchard, 1850

Distribution: Mexico, Guatemala, Belize, El Salvador, and Honduras (^{Berry, 1959b}; ^{Schoolmeesters, 2022}).

Locality records: LIBERTAD: Santa Tecla (^{Berry & Salazar-Vaquero, 1957: 47}; Bader, 1992: 327) (Fig. 17).

Years of collection: no data.

Months of collection: no data.

Topographic zone: coastal mountains.

Elevation range: 900 m.

Comments: reported as S. mexicana and Strigodermella intermedia Casey by ^{Berry and Salazar-Vaquero (1957)}, and ^{Berry (1959b)}. ^{Berry and Salazar-Vaquero (1957)} mentioned that the adults feed on rosebush. Map 2 in Bader (1992, p. 327) shows 2 points that do not correlate with the Salvadoran locality record known for the species. Strigoderma sulcipennis Burmeister, 1844

Distribution: Mexico, Guatemala, Belize, El Salvador, Honduras, Nicaragua, Costa Rica, Panama, Ecuador, Bolivia, and Argentina (^{Schoolmeesters, 2022}).

Locality records: Chalatenango: San Ignacio (1 km N Puente Julupa) (RDCC; Bader, 1992: 330); San Salvador: Parque Saburo Hirao (Río La Danta) (MUHNES; Bader, 1992: 330) (Fig. 17); no specific department: no specific locality (^{Berry, 1959b: 80}).

Years of collection: 1976-1979.

Months of collection: April and August.

Topographic zone: northern mountains, interior valley.

Elevation range: 700-1,060 m.

Paranomala Casey, 1915 Paranomala amphicoma (Bates, 1887)

Distribution: El Salvador and Panama (^{Franz, 1955}; ^{Schoolmeesters, 2022}).

Locality records: San Vicente: Volcán de San Vicente-Chinchontepec (Hacienda El Carmen) (^{Franz, 1955: 204}) (Fig. 17).

Years of collection: 1951.

Months of collection: October.

Topographic zone: coastal mountains.

Elevation range: 2,000 m.

Comments: the year of collection of the specimens is specified in ^{Hincks (1953)}.

Paranomala championi (Bates, 1887)

Distribution: Guatemala (^{Schoolmeesters, 2022}) and El Salvador.

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (RDCC) (Fig. 17).

Years of collection: 1999, 2000.

Months of collection: May.

Topographic zone: coastal mountains.

Elevation range: 780 m.

Comments: new country record.

Paranomala cincta cincta (Say, 1835)

Distribution: Mexico, Belize, Guatemala, El Salvador, and Colombia (^{Berry, 1959a}; ^{Schoolmeesters, 2022}).

Locality records: La Libertad: Santa Tecla (^{Berry & Salazar-Vaquero, 1957: 43}) (Fig. 17).

Years of collection: no data.

Months of collection: no data.

Topographic zone: coastal mountains.

Elevation range: 900 m.

Comments: reported in the literature as Anomala cincta. ^{Berry and Salazar-Vaquero (1957)} reported the adults are abundant in the first half-hour after sunset, after which they disappear. The same authors mentioned that the adults feed on rosebush and other ornamental plants. ^{Berry (1959a)} noted that host plants in El Salvador are “garden plants”.

Paranomala cupricollis (Chevrolat, 1834)

Distribution: Mexico, Guatemala, Belize, El Salvador, Honduras, Nicaragua, Costa Rica, Panama, and Venezuela (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (EAPZ, RDCC); La Libertad: Santa Tecla (^{Berry & Salazar-Vaquero, 1957: 43}); San Salvador: San Salvador (^{Franz, 1955: 204}) (Fig. 17).

Years of collection: 1951, 1999-2000.

Months of collection: May.

Topographic zone: interior valley, coastal mountains.

Elevation range: 685-900 m.

Comments: reported in the literature as Anomala cupricollis. The year of collection of the specimens from San Salvador is reported in ^{Hincks (1953)}. ^{Berry and Salazar-Vaquero (1957)}, and ^{Berry (1959a)} mentioned that the species is common in the coffee plantations of El Salvador but is not harmful to the plants.

Paranomala denticollis (Bates, 1887)

Distribution: Guatemala (^{Schoolmeesters, 2022}) and El Salvador.

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (RDCC) (Fig. 17).

Years of collection: 1999, 2000.

Months of collection: May.

Topographic zone: coastal mountains.

Elevation range: 780 m.

Comments: new country record. Paranomala donovani (Stephens, 1830)

Distribution: Mexico and El Salvador (^{Schoolmeesters, 2022}).

Locality records: San Salvador: San Salvador (^{Franz, 1955: 204}) (Fig. 17).

Years of collection: 1951.

Months of collection: June.

Topographic zone: interior valley.

Elevation range: 685 m.

Comments: the year of collection of the specimens is specified in ^{Hincks (1953)}.

Paranomala eucoma (Bates, 1887)

Distribution: Guatemala, Costa Rica (^{Schoolmeesters, 2022}), and El Salvador.

Locality records: La Libertad: Santa Tecla (EAPZ) (Fig. 18).

Years of collection: 1976.

Months of collection: August.

Topographic zone: coastal mountains.

Elevation range: 900 m.

Comments: new country record.

Paranomala flavilla (Bates, 1888)

Distribution: Mexico and El Salvador (^{Andrews et al., 1979}; ^{Schoolmeesters, 2022}).

Locality records: no specific department: no specific locality (^{Andrews et al., 1979: 7}).

Years of collection: 1978.

Months of collection: no data.

Topographic zone: no data.

Elevation range: no data.

Comments: reported as Anomala flavilla by ^{Andrews et al. (1979)}.

Paranomala foraminosa (Bates, 1887)

Distribution: Mexico, Guatemala, Belize, El Salvador, Costa Rica, and Panama (^{Schoolmeesters, 2022}).

Locality records: San Salvador: San Salvador (^{Franz, 1955: 204}); San Vicente: Volcán de San Vicente-Chinchontepec (Hacienda El Carmen) (^{Franz, 1955: 204}) (Fig. 18).

Years of collection: 1951.

Months of collection: April and June.

Topographic zone: interior valley, coastal mountains.

Elevation range: 685-2,000 m.

Comments: reported as Anomala cupricollis by ^{Franz (1955)}. The year of collection of the specimens is reported in ^{Hincks (1953)}.

Paranomala inconstans (Burmeister, 1844)

Distribution: Mexico, Guatemala, El Salvador, Honduras, Nicaragua, Costa Rica, Panama, Colombia, Brazil, Peru, St. Lucia, and St. Vincent & the Grenadines (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (EAPZ); Santa Ana: Parque Nacional Montecristo (San José Ingenio) (MUHNES); San Salvador: San Salvador (^{Franz, 1955: 204}); Cabañas: Tejutepeque (El Tamagás) (MUHNES); Morazán: Arambala (Río Sapo) (MUHNES); La Unión: La Unión (^{Berry & Salazar-Vaquero, 1957: 43}) (Fig. 18).

Years of collection: 1951, 1999-2001.

Months of collection: April and May.

Topographic zone: northern mountains, interior valley, coastal mountains, coastal plain.

Elevation range: 30-810 m.

Comments: reported as Anomala inconstans by ^{Franz (1955)}, ^{Berry (1959b)}, and ^{Saunders et al. (1998)}. Paranomala ochrogastra (Bates, 1887)

Distribution: El Salvador, Costa Rica, and Panama (^{Serrano-Chicas, 2019}; ^{Schoolmeesters, 2022}).

Locality records: Santa Ana: Parque Nacional Montecristo (Majaditas) (^{Serrano-Chicas, 2019: 41, 42}) (Fig. 18).

Years of collection: 2019.

Months of collection: April.

Topographic zone: northern mountains.

Elevation range: 1,070 m.

Paranomala pincelada (Filippini, Galante, & Micó, 2015) Distribution: El Salvador and Costa Rica (^{Serrano-Chicas, 2019}; ^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (^{Pablo-Cea, 2021: 21}); Santa Ana: El Congo (RDCC); Parque Nacional Montecristo (San José Ingenio) (^{Serrano-Chicas, 2019: 41, 42}); La Libertad: Parque El Bicentenario (^{Pablo-Cea, 2021: 21}); San Salvador: San Salvador (RDCC) (Fig. 18).

Years of collection: 1979, 2000, 2019.

Months of collection: April, May, and September-November.

Topographic zone: northern mountains, interior valley, coastal mountains.

Elevation range: 685-870 m. Paranomala plurisulcata (Bates, 1887)

Distribution: Guatemala, Nicaragua (^{Schoolmeesters, 2022}), and El Salvador.

Locality records: Usulután: no specific locality (EAPZ) (Fig. 18).

Years of collection: no data.

Months of collection: no data.

Topographic zone: no data.

Elevation range: no data.

Comments: new country record.

Paranomala quiche (Ohaus, 1897)

Distribution: Guatemala, El Salvador, and Costa Rica (^{Serrano-Chicas, 2019}; ^{Schoolmeesters, 2022}).

Locality records: Santa Ana: Parque Nacional Montecristo (Majaditas) (^{Serrano-Chicas, 2019: 41, 42}) (Fig. 18).

Years of collection: 2019.

Months of collection: March.

Topographic zone: northern mountains.

Elevation range: 1,070 m.

Paranomala robiginosa (Filippini, Galante, & Micó, 2015)

Distribution: El Salvador and Costa Rica (^{Serrano-Chicas, 2019}; ^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (^{Pablo-Cea, 2021}); Santa Ana: Parque Nacional Montecristo (La Torre, San José Ingenio) (^{Serrano-Chicas, 2019: 41, 42}) (Fig. 18).

Years of collection: 2019.

Months of collection: April and May.

Topographic zone: northern mountains, coastal mountains.

Elevation range: 780-1,450 m. Paranomala semicincta (Bates, 1887)

Distribution: Mexico, Nicaragua, Costa Rica (^{Schoolmeesters, 2022}), and El Salvador.

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (RDCC) (Fig. 18).

Years of collection: 2000.

Months of collection: May.

Topographic zone: coastal mountains.

Elevation range: 780 m.

Comments: new country record. Paranomala sticticoptera (Blanchard, 1850)

Distribution: Mexico, Guatemala, Belize, Costa Rica (^{Schoolmeesters, 2022}), and El Salvador.

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (RDCC); Santa Ana: Parque Nacional Montecristo (Los Planes) (RDCC) (Fig. 18).

Years of collection: 1999-2000.

Months of collection: May.

Topographic zone: northern mountains, coastal mountains.

Elevation range: 780-1,890 m.

Comments: new country record. Paranomala testaceipennis (Blanchard, 1850)

Distribution: El Salvador, Nicaragua, Costa Rica, Panama, Colombia, Ecuador, Bolivia, and Argentina (^{Schoolmeesters, 2022}).

Locality records: Santa Ana: Parque Nacional Montecristo (Majaditas) (^{Serrano-Chicas, 2019: 42}); La Libertad: Santa Tecla (^{Berry & Salazar-Vaquero, 1957: 43}); San Salvador: San Salvador (^{Franz, 1955: 204}) (Fig. 19).

Years of collection: 1952, 2019.

Months of collection: April, May, and November.

Topographic zone: northern mountains, interior valley, coastal mountains.

Elevation range: 685-1,070 m.

Comments:^{Berry and Salazar-Vaquero (1957)} mentioned that the species is abundant at sunset. ^{Berry (1959a)} observed that the host plants in El Salvador are “garden plants” (rosebush [Rosa spp.] and other ornamental plants).

Paranomala trapezifera (Bates, 1887)

Distribution: Mexico, El Salvador, and Costa Rica (^{Berry & Salazar-Vaquero, 1957}; ^{Schoolmeesters, 2022}).

Locality records: San Vicente: Santa Cruz Porrillo (^{Berry & Salazar-Vaquero, 1957: 43}) (Fig. 19).

Years of collection: no data.

Months of collection: no data.

Topographic zone: interior valley.

Elevation range: 40 m.

Paranomala undulata undulata (Melsheimer, 1845) Distribution: Canada, USA, Mexico, Guatemala, Costa

Rica (^{Schoolmeesters, 2022}), and El Salvador.

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (RDCC) (Fig. 19).

Years of collection: 2000.

Months of collection: May.

Topographic zone: coastal mountains.

Elevation range: 780 m.

Comments: new country record. Paranomala veraecrucis (Bates, 1887)

Distribution: Mexico, El Salvador, and Costa Rica (^{Berry & Salazar-Vaquero, 1957}; ^{Schoolmeesters, 2022}).

Locality records: La Libertad: Santa Tecla (^{Berry & Salazar-Vaquero, 1957: 43}) (Fig. 19).

Years of collection: 1951.

Months of collection: June.

Topographic zone: coastal mountains.

Elevation range: 900 m. Paranomala vicenti (^{Franz, 1955})

Distribution: El Salvador (^{Schoolmeesters, 2022}).

Locality records: San Vicente: Volcán de San Vicente-Chinchontepec (Hacienda El Carmen) (^{Franz, 1955: 205}) (Fig. 19).

Years of collection: no data.

Months of collection: coastal mountains.

Topographic zone: 2,000 m.

Comments: named after the department and Volcán San Vicente (Chinchontepec). The species is precinctive to El Salvador.

Tribe Anoplognathini MacLeay, 1819 Phalangogonia Burmeister, 1844 Phalangogonia punctata Franz, 1955

Distribution: El Salvador (^{Schoolmeesters, 2022}).

Locality records: Santa Ana: Parque Nacional Montecristo (Bosque Nebuloso) (^{Franz, 1955: 207}; ^{Smith & Morón, 2003: 333-334}) (Fig. 19). Years of collection: 1951. Months of collection: June. Topographic zone: northern mountains. Elevation range: 2,360 m.

Tribe Geniatini Burmeister, 1844 Leucothyreus MacLeay, 1819 Leucothyreus femoratus Burmeister, 1844

Distribution: Mexico, Nicaragua, El Salvador, Costa Rica, Panama, Colombia, and Venezuela (^{Schoolmeesters, 2022}).

Locality records: San Salvador: San Salvador (^{Franz, 1955: 207}; ^{Berry & Salazar-Vaquero, 1957: 45}) (Fig. 19).

Years of collection: 1951

Months of collection: February, April, and June.

Topographic zone: interior valley.

Elevation range: 685 m.

Tribe Rutelini MacLeay, 1819

Chrysina Kirby, 1828

Chrysina karschi (Nonfried, 1891)

Distribution: Guatemala, El Salvador, and Honduras (^{Schoolmeesters, 2022})

Locality records: Santa Ana: Parque Nacional Montecristo (Bosque Nebuloso) (^{López-Sorto & Sermeño-Chicas, 2013: 54}); Chalatenango: Cerro El Pital (^{López-Sorto & Sermeño-Chicas, 2013: 54}); La Montañona (^{López-Sorto & Sermeño-Chicas, 2013: 54}) (Fig. 19).

Years of collection: no data.

Months of collection: no data.

Topographic zone: northern mountains.

Elevation range: 1,600-2,660 m.

Chrysina pehlkei (Ohaus, 1930)

Distribution: Guatemala and El Salvador (^{School-meesters, 2022})

Locality records: Chalatenango: La Montañona (^{López-Sorto & Sermeño-Chicas, 2013: 54}) (Fig. 19).

Years of collection: no data.

Months of collection: no data.

Topographic zone: northern mountains.

Elevation range: 1,600 m. Chrysina quetzalcoatli (Morón, 1990)

Distribution: Mexico, Guatemala, El Salvador, Honduras, and Nicaragua (^{Schoolmeesters, 2022})

Locality records: Santa Ana: Parque Nacional Montecristo (Bosque Nebuloso) (^{López-Sorto & Sermeño-Chicas, 2013: 54}); (Los Planes) (MUHNES); Chalatenango: Cerro El Pital (MUHNES; ^{López-Sorto & Sermeño-Chicas, 2013: 54}); La Montañona (^{López-Sorto & Sermeño-Chicas, 2013: 54}) (Fig. 19). Years of collection: 1976, 1999-2002.

Months of collection: May, October, and November.

Topographic zone: northern mountains.

Elevation range: 1,600-2,660 m.

Cnemida Kirby, 1827 Cnemida aterrima Bates, 1888

Distribution: Mexico, Guatemala, El Salvador, Nicaragua, and Costa Rica (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (RDCC); La Libertad: Santa Tecla (^{Berry & Salazar-Vaquero, 1957: 44}); Cuscatlán: no specific locality (^{Jameson, 1996: 290}) (Fig. 20).

Years of collection: 1979.

Months of collection: June.

Topographic zone: coastal mountains.

Elevation range: 780-900 m.

Elcarmeniella Franz, 1955 Elcarmeniella striata Franz, 1955

Distribution: Guatemala, El Salvador, and Honduras (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (EAPZ, RDCC); San Vicente: Volcán de San Vicente-Chinchontepec (Hacienda El Carmen) (^{Franz, 1955: 203}; ^{Morón, 1987: 82}) (Fig. 20).

Years of collection: 1951, 1999.

Months of collection: May.

Topographic zone: coastal mountains.

Elevation range: 780-2,000 m.

Comments: the specimens reported by ^{Franz (1955)} were taken in the same collection event in 1951 as the passalid specimens reported by ^{Hincks (1953)}. The scientific name is based on El Carmen Farm, the type locality (^{Franz, 1955}).

Heterosternus Dupont, 1832 Heterosternus rodriguezi Candèze, 1869

Distribution: Mexico, Guatemala, and El Salvador (^{Schoolmeesters, 2022}).

Locality records: Santa Ana: Parque Nacional Montecristo (Bosque Nebuloso) (^{Morón, 1983: 40}); (Los Planes) (MUHNES) (Fig. 20).

Years of collection: 1971, 2002.

Months of collection: May and June.

Topographic zone: northern mountains.

Elevation range: 1,890-2,360 m.

Comments: the year 1971 and month of May are specified in ^{Howden & Peck (1972)}.

Macraspis MacLeay, 1819

Macraspis aterrima (Waterhouse, 1881)

Distribution: Mexico, Guatemala, El Salvador, and Honduras (^{Schoolmeesters, 2022}).

Locality records: no specific department: no specific locality (^{Morón & Paucar-Cabrera, 2003: 474}).

Years of collection: no data.

Months of collection: no data.

Topographic zone: no data.

Elevation range: no data.

Mesosternus^{Morón, 1987} Mesosternus halffteri^{Morón, 1987}

Distribution: Mexico, Guatemala, El Salvador, Honduras, and Nicaragua (^{Schoolmeesters, 2022}).

Locality records: Santa Ana: Parque Nacional Montecristo (Bosque Nebuloso) (^{Morón & Howden, 1992: 18}) (Fig. 20).

Years of collection: 1971.

Months of collection: May.

Topographic zone: northern mountains.

Elevation range: 2,360 m.

Comments: specimens were captured at blacklight in Parque Nacional Montecristo about 20:00 hrs in a light rain (^{Morón & Howden, 1992}).

Parisolea Bates, 1888

Parisolea pallida (Candèze, 1869)

Distribution: Mexico, Guatemala, El Salvador, Honduras, Nicaragua, Costa Rica, and Ecuador (^{Schoolmeesters, 2022}).

Locality records: Santa Ana: Parque Nacional Montecristo (Los Planes) (^{Delgado et al., 2006: 95}); San Salvador: San Salvador (^{Delgado et al., 2006: 95}) (Fig. 20).

Years of collection: 1991.

Months of collection: May.

Topographic zone: northern mountains, interior valley.

Elevation range: 685-1,890 m. Parisolea pachytarsis (^{Morón, 1987})

Distribution: Mexico, Guatemala, El Salvador, Honduras, Nicaragua, and Costa Rica (^{Schoolmeesters, 2022}).

Locality records: Santa Ana: Parque Nacional Montecristo (^{Morón & Howden, 1992: 18-19}) (Bosque Nebuloso, La Torre, Los Planes, Majaditas) (^{Serrano-Chicas, 2019: 42}); San Salvador: San Salvador (^{Morón, 1987: 73}) (Fig. 20).

Years of collection: 1953, 1971, 1999, 2019.

Months of collection: March-May.

Topographic zone: northern mountains, interior valley.

Elevation range: 685-2,360 m.

Comments: specimens were captured at a blacklight in Parque Nacional Montecristo about 20:00 hrs in a light rain (^{Morón & Howden, 1992}) and at a mercury vapor lamp (R. D. Cave, personal observation).

Pelidnota MacLeay, 1819

Pelidnota guatemalensis Bates, 1888

Distribution: Mexico, Guatemala, Belize, Honduras (^{Schoolmeesters, 2022}), and El Salvador.

Locality records: Sonsonate: Izalco (Los Guates Farm) (MUHNES); Cabañas: Tejutepeque (El Tamagás) (MUHNES); San Miguel: Laguna El Jocotal (MUHNES); La Unión: Isla Martín Pérez (MUHNES) (Fig. 20).

Years of collection: 1974, 2000-2001.

Months of collection: May, August, September, and November.

Topographic zone: interior valley, coastal plain.

Elevation range: 25-700 m.

Comments: new country record.

Pelidnota notata Blanchard, 1851

Distribution: Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Colombia, Venezuela, Ecuador (^{Schoolmeesters, 2022}), and El Salvador.

Locality records: San Salvador: Ruinas de Cihuatán (MUHNES) (Fig. 20).

Years of collection: 1978.

Months of collection: September.

Topographic zone: interior Valley.

Elevation range: 325 m.

Comments: new country record.

Pelidnota punctulata Bates, 1888

Distribution: Mexico, Guatemala, Belize, El Salvador, Honduras, Nicaragua, Costa Rica, Panama, Colombia, Venezuela, and Ecuador (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (EAPZ, RDCC); (San Francisco Menéndez) (MUHNES); Santa Ana: Parque Nacional Montecristo (San José Ingenio) (MUHNES); La Libertad: Los Chorros (RDCC); Quezaltepeque (^{Hardy, 1975: 21}; ^{Moore et al., 2017: 246}); San Salvador: San Salvador (^{Franz, 1955: 203}; ^{Hardy, 1975: 21}; ^{Moore et al., 2017: 246}); Cuscatlán: San Cristóbal (^{Hardy, 1975: 21}); Usulután: Nancuchiname (MUHNES); Morazán: Cerro Cacahuatique (MUHNES) (Fig. 20); no specific department: no specific locality (^{Berry, 1959b: 60}).

Years of collection: 1951-1952, 1977-1979, 1999-2001.

Months of collection: April-September.

Topographic zone: northern mountains, interior valley, coastal mountains, coastal plain.

Elevation range: 15-1,400 m.

Comments: the specimens reported by ^{Franz (1955)} were taken in the same collection event in 1951 as the passalid specimens reported by ^{Hincks (1953)}.

Pelidnota strigosa Laporte, 1840

Distribution: Mexico, Guatemala, Belize, El Salvador,

Honduras, Nicaragua, Costa Rica, Panama, Colombia, and Venezuela (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (J. Pablo, personal observation); Santa Ana: Lago de Coatepeque (^{Hardy, 1975: 20}; ^{Moore et al., 2017: 263}); Flor Amarilla (Abajo) (^{Berry & Salazar-Vaquero, 1957: 46}; ^{Hardy, 1975: 20}; ^{Moore et al., 2017: 263}); Parque Nacional Montecristo (MUHNES); (San José Ingenio) (^{Serrano-Chicas, 2019: 42}); La Libertad: Los Chorros (RDCC); Quezaltepeque (^{Hardy, 1975: 20}; ^{Moore et al., 2017: 263}); Santa Tecla (^{Franz, 1955: 203}); San Salvador: Lago de Ilopango (^{Hardy, 1975: 20}; ^{Moore et al., 2017: 263}); San Salvador (^{Franz, 1955: 204}); Cuscatlán: San Cristóbal (Border Station) (^{Hardy, 1975: 20}; ^{Moore et al., 2017: 263}); La Unión: La Unión (^{Hardy, 1975: 20}; ^{Moore et al., 2017: 263}) (Fig. 20).

Years of collection: 1950, 1976-1979, 2013, 2019.

Months of collection: April, and July-December.

Topographic zone: northern mountains, interior valley, coastal mountains, coastal plain.

Elevation range: 30-900 m.

Comments: the specimens reported by ^{Franz (1955)} were taken in the same collection event in 1951 as the passalid specimens reported by ^{Hincks (1953)}. ^{Hardy (1975)} reported examining material from “Flor Amaysta Ana”. It is assumed that the place is Flor Amarilla (Abajo), in Santa Ana (“...sta Ana” in abbreviation). ^{Berry & Salazar (1957)} reported the species in the same place. ^{Hardy (1975)} also reported specimens collected at “Lunada Beach”, a locality unknown to us. ^{Berry & Salazar (1957)} and ^{Berry (1959a)} mentioned that the species is common in coffee plantations of El Salvador but is not harmful to the plants. Pelidnota virescens Burmeister, 1844

Distribution: Mexico, El Salvador, Honduras, Nicaragua, and Costa Rica (^{Berry 1959a}; ^{Schoolmeesters, 2022}).

Locality records: Santa Ana: Flor Amarilla (Abajo) (^{Berry & Salazar-Vaquero, 1957: 46}); La Libertad: San Andrés (^{Berry & Salazar-Vaquero, 1957: 46}); Santa Tecla (^{Berry & Salazar-Vaquero, 1957: 46}) (Fig. 21).

Years of collection: no data.

Months of collection: no data.

Topographic zone: interior valley, coastal mountains.

Elevation range: 475-900 m.

Comments:^{Berry and Salazar-Vaquero (1957)}, and ^{Berry (1959a)} mentioned that the species is common in cultivated coffee and corn in El Salvador but is not harmful to the plants.

Subfamily Dynastinae MacLeay, 1819 Tribe Agaocephalini Burmeister, 1847 Spodistes Burmeister, 1847 Spodistes mniszechi (Thomson, 1860)

Distribution: Mexico, Guatemala, Belize, El Salvador, Honduras, Nicaragua, Costa Rica, Panama, and Colombia (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (^{Ratcliffe & Cave, 2006: 353}); Santa Ana: Parque Nacional Montecristo (Bosque Nebuloso) (MUHNES); (Los Planes) (^{Ratcliffe & Cave, 2006: 354}); Chalatenango: La Montañona (^{Ratcliffe & Cave, 2006: 353}); San Ignacio (MUHNES); La Libertad: Los Chorros (^{Ratcliffe & Cave, 2006: 353}); San Andrés (^{Ratcliffe & Cave, 2006: 353}); Santa Tecla (^{Ratcliffe & Cave, 2006: 353}); San Salvador: Cerro San Jacinto (^{Ratcliffe & Cave, 2006: 354}); San Salvador (^{Ratcliffe & Cave, 2006: 354}); La Paz: Santiago Nonualco (^{Ratcliffe & Cave, 2006: 353}); Cabañas: Cinquera (^{Ratcliffe & Cave, 2006: 353}) (Fig. 21).

Years of collection: 1959, 1976, 1997-2002.

Months of collection: All year.

Topographic zone: northern mountains, interior valley, coastal mountains, coastal plain.

Elevation range: 160-2,360 m. Spodistes monzoni Warner, 1992

Distribution: Mexico, Guatemala, and El Salvador (^{Schoolmeesters, 2022}).

Locality records: Santa Ana: Parque Nacional Montecristo [probably San José Ingenio] (^{Ratcliffe & Cave, 2006: 355}; ^{Serrano-Chicas, 2019: 37, 38}); San Salvador: San Salvador (^{Warner, 1992: 378}; ^{Ratcliffe & Cave, 2006: 355}); Morazán: Arambala (Río Sapo) (^{Ratcliffe & Cave, 2006: 355}); La Unión: Volcán de Conchagua (^{Ratcliffe & Cave, 2006: 355}) (Fig. 21).

Years of collection: 1960, 1997-2002.

Months of collection: April and May.

Topographic zone: northern mountains, interior valley, coastal mountains.

Elevation range: 665-1,190 m.

Tribe Cyclocephalini Laporte, 1840 Ancognatha Erichson, 1847 Ancognatha sellata Arrow, 1911

Distribution: Mexico, Guatemala, El Salvador, Honduras, and Nicaragua (^{Ratcliffe & Cave, 2006}; ^{Schoolmeesters, 2022}).

Locality records: Santa Ana: Parque Nacional Cerro Verde (^{Ratcliffe & Cave, 2006: 49}); Parque Nacional Montecristo (Bosque Nebuloso) (^{Ratcliffe & Cave, 2006: 49}; ^{Serrano-Chicas, 2019: 37, 38}) (Los Planes) (^{Ratcliffe & Cave, 2006: 49}); Chalatenango: Cerro El Pital (Ratcliffe & Cave, 2006: 49); San Salvador: Hacienda Los Planes (UNSM) (Fig. 21).

Years of collection: 1972-1975, 1997-2002, 2019.

Months of collection: March-October, and December.

Topographic zone: northern mountains, interior valley, coastal mountains.

Elevation range: 720-2,660 m.

Aspidolea Bates, 1888

Aspidolea fuliginea (Burmeister 1847)

Distribution: Mexico, Guatemala, El Salvador, Honduras, Nicaragua, Costa Rica, Panama, Colombia, Venezuela, Trinidad & Tobago, Ecuador, Brazil, Peru, and Argentina (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (La Fincona) (^{Ratcliffe & Cave, 2006: 53}); (San Benito) (^{Ratcliffe & Cave, 2006: 53}); Santa Ana: Parque Nacional Montecristo (Majaditas, San José Ingenio) (^{Serrano-Chicas, 2019: 37, 38}); La Libertad: Santa Tecla (^{Ratcliffe & Cave, 2006: 53}); Cuscatlán: Cojutepeque (^{Ratcliffe & Cave, 2006: 53}); San José Guayabal (^{Ratcliffe & Cave, 2006: 53}) (Fig. 21).

Years of collection: 1997-2002, 2018.

Months of collection: All year.

Topographic zone: northern mountains, interior valley, coastal mountains.

Elevation range: 560-1,070 m.

Aspidolea singularis Bates, 1888

Distribution: USA, Mexico, Guatemala, El Salvador, Honduras, Nicaragua, Costa Rica, Panama, Colombia, Venezuela, Ecuador, Brazil, and Peru (^{Schoolmeesters, 2022})

Locality records: Ahuachapán: Parque Nacional El Imposible (La Fincona) (^{Ratcliffe & Cave, 2006: 54}); (San Benito) (^{Ratcliffe & Cave, 2006: 54}); Santa Ana: Parque Nacional Montecristo (Los Planes) (^{Ratcliffe & Cave, 2006: 54}; ^{Serrano-Chicas, 2019: 37, 38}); La Libertad: Los Chorros (^{Ratcliffe & Cave, 2006: 54}); Parque El Bicentenario (^{Pablo-Cea, 2021: 21}; ^{Pablo-Cea, Deloya, MacGregor-Fors, & Navarrete-Heredia, 2022: 277}; ^{Pablo-Cea, Deloya, MacGregor-Fors, Navarrete-Heredia et al., 2022: 378}); Santa Tecla (^{Ratcliffe & Cave, 2006: 54}); Zapotitán (^{Ratcliffe & Cave, 2006: 54}); San Salvador: San Salvador (^{Ratcliffe & Cave, 2006: 54}); Zacamil (^{Ratcliffe & Cave, 2006: 54}); Morazán: Arambala (Río Sapo) (^{Ratcliffe & Cave, 2006: 54}) (Fig. 21).

Years of collection: 1961, 1997-2002, 2019.

Months of collection: All year.

Topographic zone: northern mountains, interior valley, coastal mountains.

Elevation range: 480-1,890 m.

Comments:^{Ratcliffe and Cave (2006)} reported specimens collected in San Antonio Chávez in San Salvador, but this locality is unknown to us.

Cyclocephala Dejean, 1821 Cyclocephala amblyopsis Bates, 1888

Distribution: Mexico, Guatemala, El Salvador, Honduras, Nicaragua, Costa Rica, Panama, Colombia, Ecuador, Brazil, Peru, and Bolivia (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (^{Ratcliffe & Cave, 2006: 74}); Santa Ana: Parque Nacional Montecristo (Majaditas, San José Ingenio) (^{Serrano-Chicas, 2019: 37, 38}); La Libertad: Puerto de La Libertad (^{Romero-Nápoles, 2021}) (Fig. 21).

Years of collection: 1997-2002, 2018.

Months of collection: January, and March-November.

Topographic zone: northern mountains, coastal mountains, coastal plain.

Elevation range: 30-1,070 m. Cyclocephala complanata Burmeister, 1847

Distribution: Mexico, Guatemala, El Salvador, Honduras, Nicaragua, Costa Rica, and Panama (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (^{Ratcliffe & Cave, 2006: 89}); Santa Ana: Parque Nacional Montecristo (La Torre, Majaditas) (^{Serrano-Chicas, 2019: 37, 38}); (Los Planes) (^{Ratcliffe & Cave, 2006: 89}); Chalatenango: San Ignacio (^{Ratcliffe & Cave, 2006: 89}) (Fig. 21); no specific department: no specific locality (^{Ratcliffe & Morón, 1997: 59}).

Years of collection: 1979, 1997-2002, 2019.

Months of collection: April-September.

Topographic zone: northern mountains, coastal mountains.

Elevation range: 780-1,890 m. Cyclocephala confusa Endrödi, 1966

Distribution: Mexico, Guatemala, Belize, El Salvador, Honduras, Nicaragua, Costa Rica, Panama, Colombia, and Peru (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (^{Ratcliffe & Cave, 2006: 94}); Santa Ana: Parque Nacional Montecristo (Majaditas) (^{Serrano-Chicas, 2019: 37, 38}); Morazán: Arambala (Río Sapo) (^{Ratcliffe & Cave, 2006: 94}) (Fig. 21).

Years of collection: 1997-2002, 2019.

Months of collection: April-September.

Topographic zone: northern mountains, coastal mountains.

Elevation range: 665-1,070 m.

Comments: reported as Cyclocephala confunsa [sic] by ^{Serrano-Chicas (2019)}.

Cyclocephala curta Bates, 1888

Distribution: Mexico, El Salvador, Honduras, Nicaragua, and Costa Rica (^{Schoolmeesters, 2022})

Locality records: no specific department: no specific locality (^{Ratcliffe & Cave, 2006: 99}).

Years of collection: 1999.

Months of collection: March.

Topographic zone: no data.

Elevation range: no data. Cyclocephala deceptor (Casey, 1915)

Distribution: Mexico, El Salvador, Honduras, and Nicaragua (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (^{Ratcliffe & Cave, 2006: 100}; ^{Pablo-Cea, 2021: 21}; ^{Pablo-Cea, Deloya, MacGregor-Fors, Navarrete-Heredia, & Espinosa-de los Monteros, 2022: 378}); Santa Ana: Coatepeque (^{Ratcliffe & Cave, 2006: 100}); Metapán (^{Ratcliffe & Cave, 2006: 100}); Parque Nacional Montecristo (Majaditas, San José Ingenio) (^{Serrano-Chicas, 2019: 37, 38}); (Los Planes) (^{Ratcliffe & Cave, 2006: 100}); Sonsonate: Volcán de Izalco (^{Ratcliffe & Cave, 2006: 100}); Chalatenango: San Ignacio (MUHNES); La Libertad: La Libertad (^{Ratcliffe & Cave, 2006: 100}); Los Chorros (^{Ratcliffe & Cave, 2006: 100}); San Andrés (^{Ratcliffe & Cave, 2006: 100}); Santa Tecla (^{Ratcliffe & Cave, 2006: 100}); San Salvador: Aguilares (^{Ratcliffe & Cave, 2006: 100}); Cerro San Jacinto (^{Ratcliffe & Cave, 2006: 100}); Puerta del Diablo (^{Ratcliffe & Cave, 2006: 100}); San Salvador (Alta María, Los Planes) (^{Ratcliffe & Cave, 2006: 100}); Tonacatepeque (^{Ratcliffe & Cave, 2006: 100}); La Paz: San Luis [Talpa] (^{Ratcliffe & Cave, 2006: 100}); San Vicente: San Cayetano Istepeque (^{Ratcliffe & Cave, 2006: 100}); Morazán: Arambala (Río Sapo) (^{Ratcliffe & Cave, 2006: 100}); no specific department: no specific locality (^{Berry, 1959b: 79}) (Fig. 21).

Years of collection: 1957-1958, 1964, 1970-1979, 1997-2002, 2019.

Months of collection: January-November.

Topographic zone: northern mountains, interior valley, coastal mountains, coastal plain.

Elevation range: 40-1,890 m.

Comments: reported as Stigmalia cuernavacana Casey by ^{Berry (1959b)}.

Cyclocephala freudei Endrödi, 1963

Distribution: USA, Mexico, Guatemala, El Salvador, Costa Rica, and Ecuador (^{Schoolmeesters, 2022}).

Locality records: San Salvador: San Salvador (^{Ratcliffe & Cave, 2006: 108}); Cuscatlán: El Rosario (^{Ratcliffe & Cave, 2006: 108}) (Fig. 22).

Years of collection: 1959, 1997-2002.

Months of collection: June and July.

Topographic zone: interior valley.

Elevation range: 685-720 m. Cyclocephala gravis Bates, 1888

Distribution: Mexico, Guatemala, El Salvador, Honduras, Nicaragua, Costa Rica, Panama, Colombia, Venezuela, Guyana, Ecuador, Brazil, and Bolivia (^{Schoolmeesters, 2022}).

Locality records: Morazán: Arambala (Río Sapo)

(^{Ratcliffe & Cave, 2006: 111}); Cerro Cacahuatique

(^{Ratcliffe & Cave, 2006: 111}) (Fig. 22).

Years of collection: 1997-2002.

Months of collection: January, March-October, and December.

Topographic zone: northern mountains.

Elevation range: 665-1,400 m. Cyclocephala lunulata Burmeister, 1847

Distribution: USA, Mexico, Guatemala, El Salvador, Honduras, Nicaragua, Costa Rica, Panama, Colombia, Venezuela, Trinidad & Tobago, Guyana, Ecuador, Brazil, Peru, Bolivia, Paraguay, and Argentina (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (La Fincona, San Benito) (^{Ratcliffe & Cave, 2006: 122}); Santa Ana: Parque Nacional Montecristo [probably San José Ingenio] (^{Ratcliffe & Cave, 2006: 122}; ^{Serrano-Chicas, 2019: 37, 38}); Santa Ana (^{Ratcliffe & Cave, 2006: 122}); La Libertad: Parque El Bicentenario (^{Pablo-Cea, 2021: 21}; ^{Pablo-Cea, Deloya, MacGregor-Fors, & Navarrete-Heredia, 2022: 277}; ^{Pablo-Cea, Deloya, MacGregor-Fors, Navarrete-Heredia et al., 2022: 378}); Parque Nacional Walter Thilo Deininger (^{Ratcliffe & Cave, 2006: 122}); San Andrés (^{Ratcliffe & Cave, 2006: 122}); Zapotitán (^{Ratcliffe & Cave, 2006: 122}); San Salvador: Cuscatancingo (^{Berry & Salazar-Vaquero, 1957: 44}; ^{Ratcliffe & Cave, 2006: 122}); Delgado (^{Ratcliffe & Cave, 2006: 122}); San Salvador (^{Ratcliffe & Cave, 2006: 122}); Cabañas: Cinquera (^{Ratcliffe & Cave, 2006: 122}); Tejutepeque (^{Ratcliffe & Cave, 2006: 122}); Usulután: Cerro El Tigre (^{Ratcliffe & Cave, 2006: 122}); San Miguel: Laguna El Jocotal (^{Ratcliffe & Cave, 2006: 122}); Morazán: Arambala (Río Sapo) (^{Ratcliffe & Cave, 2006: 122}); Cerro Cacahuatique (^{Ratcliffe & Cave, 2006: 122}); La Unión: Playa El Icacal (^{Ratcliffe & Cave, 2006: 122}); Volcán de Conchagua (^{Ratcliffe & Cave, 2006: 122}) (Fig. 22).

Years of collection: 1971-1978, 1997-2002, 2018-2019.

Months of collection: All year.

Topographic zone: northern mountains, interior valley, coastal mountains, coastal plain.

Elevation range: 15-1,580 m.

Comments:^{Berry and Salazar-Vaquero (1957)} mentioned that adults perforate orange fruits (Citrus sp.), which allows fungi to enter. The species is considered a part of the “gallina ciega” complex in cultivated corn in El Salvador (^{Berry, 1959b}; ^{Andrews et al., 1979}; Mendoza, 1996).

Cyclocephala mafaffa Burmeister, 1847

Distribution: USA, Mexico, Guatemala, El Salvador, Honduras, Nicaragua, Costa Rica, Panama, Colombia, Venezuela, Trinidad & Tobago, Ecuador, Brazil, and Guadeloupe (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (^{Ratcliffe & Cave, 2006: 127}; ^{Pablo-Cea, 2021: 21}; ^{Pablo-Cea, Deloya, MacGregor-Fors, Navarrete-Heredia, & Espinosa-de los Monteros, 2022: 378}); Santa Ana: Parque Nacional Montecristo (Los Planes) (^{Ratcliffe & Cave, 2006: 127}); (San José Ingenio) (^{Serrano-Chicas, 2019: 37, 38}); Santa Ana (^{Ratcliffe & Cave, 2006: 127}); Chalatenango: La Montañona (^{Ratcliffe & Cave, 2006: 127}); San Salvador: San Salvador (^{Berry &Salazar-Vaquero, 1957: 44}); Usulután: Cerro El Tigre (UNSM; ^{Ratcliffe & Cave, 2006: 127}); Usulután (^{Ratcliffe & Cave, 2006: 127}); Morazán: Arambala (Río Sapo) (^{Ratcliffe & Cave, 2006: 127}); Cerro Cacahuatique (^{Ratcliffe & Cave, 2006: 127}) (Fig. 22).

Years of collection: 1972-1979, 1997-2002, 2019.

Months of collection: All year.

Topographic zone: northern mountains, interior valley, coastal mountains, coastal plain.

Elevation range: 80-1,890 m.

Comments: reported as Cyclocephala mefaffa [sic] in ^{Berry and Salazar-Vaquero (1957)}.

Cyclocephala melanocephala (Fabricius, 1775) Distribution: USA, Mexico, Guatemala, Belize, El Salvador, Honduras, Nicaragua, Costa Rica, Panama, Colombia, Venezuela, Trinidad & Tobago, Guyana, Suriname, French Guiana, Ecuador, Brazil, Peru, Bolivia, Paraguay, Argentina, Dominican Republic, Puerto Rico, St. Kitts & Nevis, Montserrat, Guadeloupe, Dominica, Martinique, St. Lucia, St. Vincent & the Grenadines, and Grenada (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (La Fincona) (^{Ratcliffe & Cave, 2006: 130}); (San Benito) (^{Ratcliffe & Cave, 2006: 130}); Santa Ana: Parque Nacional Montecristo (Los Planes) (^{Ratcliffe & Cave, 2006: 130}); Chalatenango: La Montañona (^{Ratcliffe & Cave, 2006: 130}); La Libertad: La Libertad (CMNC); Los Chorros (CMNC); Parque Nacional Walter Thilo Deininger (^{Ratcliffe & Cave, 2006: 130}); Río Agua Caliente (^{Ratcliffe & Cave, 2006: 130}); Santa Tecla (^{Ratcliffe & Cave, 2006: 130}); San Salvador: San Salvador (CMNC); Morazán: Arambala (Río Sapo) (^{Ratcliffe & Cave, 2006: 130}); Cerro Cacahuatique (^{Ratcliffe & Cave, 2006: 130}) (Fig. 22).

Years of collection: 1959-1960, 1971-1979, 1997-2002.

Months of collection: January-November.

Topographic zone: northern mountains, interior valley, coastal mountains, coastal plain.

Elevation range: 70-1,890 m.

Cyclocephala melolonthida Ratcliffe & Cave, 2002

Distribution: Guatemala and El Salvador (^{Schoolmeesters, 2022}).

Locality records: La Paz: Playa Costa del Sol (^{Ratcliffe & Cave, 2002: 153}, ^{2006: 132}) (Fig. 22). Years of collection: 1976.

Months of collection: November.

Topographic zone: coastal plain.

Elevation range: 5 m.

Cyclocephala multiplex Casey, 1915

Distribution: Mexico, Guatemala, Belize, El Salvador, Honduras, Nicaragua, Costa Rica, and Panama (^{Schoolmeesters, 2022}).

Locality records: Santa Ana: Santa Ana (^{Ratcliffe & Cave, 2006: 134}); La Libertad: El Barillo (^{Ratcliffe & Cave, 2006: 134}); Santa Tecla (^{Ratcliffe & Cave, 2006: 134}); San Salvador: San Antonio Abad (^{Ratcliffe & Cave, 2006: 134}); San Salvador (^{Ratcliffe & Cave, 2006: 134}); Morazán: Arambala (Río Sapo) (^{Ratcliffe & Cave, 2006: 134}) (Fig. 22).

Years of collection: 1972, 1976-1979, 1997-2002.

Months of collection: All year.

Topographic zone: northern mountains, interior valley, coastal mountains.

Elevation range: 665-900 m. Cyclocephala ovulum Bates, 1888

Distribution: Mexico, Guatemala, El Salvador, Honduras, Nicaragua, Costa Rica, Panama, Colombia, Guyana, Suriname, French Guiana, Ecuador, Brazil, Paraguay, and Argentina (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (^{Ratcliffe & Cave, 2006: 140}); La Libertad: Santa Tecla (^{Ratcliffe & Cave, 2006: 140}); Zapotitán (^{Ratcliffe & Cave, 2006: 140}); San Miguel: Laguna El Jocotal (^{Ratcliffe & Cave, 2006: 140}); Morazán: Arambala (Río Sapo) (^{Ratcliffe & Cave, 2006: 140}) (Fig. 22).

Years of collection: 1997-2002.

Months of collection: All year, except January and March.

Topographic zone: northern mountains, interior valley, coastal mountains, coastal plain.

Elevation range: 25-900 m. Cyclocephala sexpunctata Laporte, 1840

Distribution: Mexico, Guatemala, El Salvador, Honduras, Nicaragua, Costa Rica, Panama, Colombia, Venezuela, French Guiana, Ecuador, Brazil, Peru, and Bolivia (^{Ratcliffe & Cave, 2006}; ^{Schoolmeesters, 2022}).

Locality records: Morazán: Cerro Cacahuatique

(^{Ratcliffe & Cave, 2006: 154}) (Fig. 22).

Years of collection: 2001.

Months of collection: July.

Topographic zone: northern mountains.

Elevation range: 1,400 m.

Comments: although relatively common in Honduras and Nicaragua, only 1 specimen from El Salvador was reported by ^{Ratcliffe and Cave (2006)}.

Cyclocephala sparsa Arrow, 1902

Distribution: Mexico, Guatemala, Belize, El Salvador, Honduras, Costa Rica, and Panama (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (^{Ratcliffe & Cave, 2006: 158}); San Salvador: San Salvador (^{Ratcliffe & Cave, 2006: 158}); Cabañas: Cinquera (^{Ratcliffe & Cave, 2006: 158}); Morazán: Cerro Cacahuatique (^{Ratcliffe & Cave, 2006: 158}) (Fig. 22).

Years of collection: 1997-2002.

Months of collection: January, April-September, and December.

Topographic zone: northern mountains, interior valley, coastal mountains.

Elevation range: 385-1,400 m. Cyclocephala stictica Burmeister, 1847

Distribution: Mexico, Guatemala, Belize, El Salvador, Honduras, Nicaragua, Costa Rica, Panama, Colombia, Venezuela, French Guiana, Ecuador, Brazil, Peru, and Bolivia (^{Schoolmeesters, 2022}).

Locality records: Santa Ana: Parque Nacional

Montecristo (Majaditas) (^{Serrano-Chicas, 2019: 37, 38});

Chalatenango: La Montañona (^{Ratcliffe & Cave, 2006: 162}); La Libertad: Parque Nacional Walter Thilo Deininger

(^{Ratcliffe & Cave, 2006: 162}); Cabañas: Tejutepeque

(^{Ratcliffe & Cave, 2006: 162}); Morazán: Arambala (Río Sapo) (^{Ratcliffe & Cave, 2006: 162}) (Fig. 23).

Years of collection: 1997-2002, 2019.

Months of collection: January, and March-November.

Topographic zone: northern mountains, interior valley, coastal mountains, coastal plain.

Elevation range: 135-1,600 m.

Cyclocephala weidneri Endrödi, 1964

Distribution: Mexico, Guatemala, El Salvador, Honduras, Costa Rica, Panama, Colombia, Venezuela, Ecuador, Brazil, Peru, and Bolivia (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (^{Ratcliffe & Cave, 2006: 167}) (Fig. 23).

Years of collection: 1976.

Months of collection: June.

Topographic zone: coastal mountains.

Elevation range: 780 m.

Comments: although relatively common in Honduras and Nicaragua, only 3 specimens from El Salvador were reported by ^{Ratcliffe and Cave (2006)}.

Dyscinetus Harold, 1869 Dyscinetus dubius (Olivier, 1789)

Distribution: Mexico, Guatemala, Belize, El Salvador, Honduras, Nicaragua, Costa Rica, Panama, Colombia, Venezuela, Trinidad & Tobago, Guyana, Suriname, French Guiana, Ecuador, Brazil, Peru, Bolivia, Paraguay, Argentina, and Cuba (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Ahuachapán (^{Ratcliffe & Cave, 2006: 171}); Parque Nacional El Imposible (San Benito) (^{Ratcliffe & Cave, 2006: 171}); San Miguelito (MUHNES); Zanjón El Chino (^{Ratcliffe & Cave, 2006: 171}); Santa Ana: Metapán (^{Ratcliffe & Cave, 2006: 171}); Parque Nacional Güija (^{Ratcliffe & Cave, 2006: 171}); Parque Nacional Los Andes (^{Ratcliffe & Cave, 2006: 171}); La Libertad: El Barillo (MUHNES); Zapotitán (^{Ratcliffe & Cave, 2006: 171}); San Salvador: San Salvador (^{Ratcliffe & Cave, 2006: 171}); Cuscatlán: Suchitoto (^{Ratcliffe & Cave, 2006: 171}); Cabañas: Cinquera (^{Ratcliffe & Cave, 2006: 171}); San Miguel: Laguna El Jocotal (^{Ratcliffe & Cave, 2006: 171}) (Fig. 23).

Years of collection: 1963-1964, 1973-1977, 1986, 1994, 1997-2002.

Months of collection: All year.

Topographic zone: interior valley, coastal mountains, coastal plain.

Elevation range: 5-805 m. Dyscinetus laevipunctatus Bates, 1888

Distribution: Mexico, Guatemala, Belize, El Salvador, Honduras, Nicaragua, Costa Rica, Panama, Colombia, and Brazil (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (^{Ratcliffe & Cave, 2006: 173}); Zanjón El Chino (^{Ratcliffe & Cave, 2006: 173}); La Libertad: San Andrés (^{Ratcliffe & Cave, 2006: 173}); San Salvador: San Salvador (^{Ratcliffe & Cave, 2006: 173}) (Fig. 23).

Years of collection: 1950, 1976-1979, 1997-2002.

Months of collection: All year, except September.

Topographic zone: interior valley, coastal mountains, coastal plain.

Elevation range: 5-1,730 m.

Stenocrates Burmeister, 1847 Stenocrates bicarinatus Robinson, 1947

Distribution: Mexico, Guatemala, Belize, El Salvador, Honduras, Nicaragua, Costa Rica, Panama, Colombia, Suriname, French Guiana, Brazil, and Peru (^{Schoolmeesters, 2022}).

Locality records: San Salvador: San Salvador (^{Ratcliffe & Cave, 2006: 181}) (Fig. 23). Years of collection: 1963.

Months of collection: October.

Topographic zone: interior valley.

Elevation range: 685 m.

Comments: although relatively common in Honduras and Nicaragua, only 1 specimen from El Salvador was reported by ^{Ratcliffe and Cave (2006)}.

Stenocrates canuli Delgado, 1991

Distribution: Mexico, Guatemala, Belize, El Salvador, Honduras, and Nicaragua (^{Schoolmeesters, 2022}).

Locality records: San Salvador: San Salvador (^{Ratcliffe & Cave, 2006: 182}) (Fig. 23). Years of collection: no data. Months of collection: no data. Topographic zone: interior valley. Elevation range: 685 m.

Comments: although relatively common in Honduras, only 1 specimen from El Salvador was reported by ^{Ratcliffe and Cave (2006)}, with no specified year and month of collection.

Tribe Dynastini MacLeay, 1819 Dynastes MacLeay, 1819 Dynastes maya Hardy, 2003

Distribution: Mexico, Guatemala, El Salvador, and Honduras (^{López-Sorto et al., 2014}; ^{Schoolmeesters, 2022}).

Locality records: Chalatenango: Cerro Malcotal (^{López-Sorto et al., 2014: 43}) (Fig. 23).

Years of collection: 2011.

Months of collection: June.

Topographic zone: northern mountains.

Elevation range: 2,190 m.

Comments: 3 adult specimens were collected as they apparently fed on decaying wood (^{López-Sorto et al., 2014}).

Golofa Hope, 1837

Golofa (Mixigenus) tersander Burmeister, 1847 Distribution: Mexico, Guatemala, El Salvador,

Honduras, Nicaragua, and Costa Rica (^{Schoolmeesters, 2022}).

Locality records: La Libertad: San Andrés (^{Ratcliffe & Cave, 2006: 380}) (Fig. 23). Years of collection: no data. Months of collection: no data. Topographic zone: interior valley. Elevation range: 475 m.

Comments: only 1 specimen is known from El Salvador, with no specified year and month of collection (^{Ratcliffe & Cave, 2006}).

Golofa (Golofa) pizarro Hope, 1837

Distribution: Mexico, Guatemala, El Salvador, Honduras, Nicaragua, Costa Rica, and Panama (^{Schoolmeesters, 2022}).

Locality records: Santa Ana: Parque Nacional Los Andes (^{Ratcliffe & Cave, 2006: 374}); Parque Nacional Montecristo (^{Ratcliffe & Cave, 2006: 374}) (Los Planes) (MUHNES); Chalatenango: Las Pilas (^{Ratcliffe & Cave, 2006: 374}); La Libertad: San Juan Opico (^{Ratcliffe & Cave, 2006: 374}) (Fig. 23).

Years of collection: 1975-1979, 1988, 1999.

Months of collection: July-September.

Topographic zone: northern mountains, interior valley, coastal mountains.

Elevation range: 480-1,890 m.

Comments: although relatively common in Honduras, only 7 specimens from El Salvador were reported by ^{Ratcliffe and Cave (2006)}.

Megasoma Kirby, 1825

Megasoma elephas (Fabricius, 1775)

Distribution: Mexico, Guatemala, Belize, El Salvador, Honduras, Nicaragua, Costa Rica, Panama, and Venezuela (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Concepción de Ataco (MUHNES); Parque Nacional El Imposible (Cerro Campana) (^{Ratcliffe & Cave, 2006: 381}); (San Benito) (^{Pablo-Cea, 2021: 22}; ^{Pablo-Cea, Deloya, MacGregor-Fors, Navarrete-Heredia, & Espinosa-de los Monteros, 2022: 379}) (San Francisco Menéndez) (^{Ratcliffe & Cave, 2006: 381}); Santa Ana: Parque Nacional Los Andés (^{Ratcliffe & Cave, 2006: 381}); Parque Nacional Montecristo (Los Planes) (^{Ratcliffe & Cave, 2006: 381}); Sonsonate: Sonsonate (MUHNES); La Libertad:

San Andrés (^{Ratcliffe & Cave, 2006: 381}); Santa Tecla (^{Ratcliffe & Cave, 2006: 381}); San Salvador: San Salvador (^{Ratcliffe & Cave, 2006: 381}); San Vicente: San Vicente (^{Ratcliffe & Cave, 2006: 381}); La Unión: Isla Zacatillo (^{Ratcliffe & Cave, 2006: 381}); no specific department: no specific locality (^{Berry, 1959b: 49}) (Fig. 23).

Years of collection: 1969, 1973-1975, 1997-2002.

Months of collection: All year.

Topographic zone: northern mountains, interior valley, coastal mountains, coastal plain.

Elevation range: 110-1,890 m.

Tribe Oryctini Mulsant, 1842

Coelosis Hope, 1837

Coelosis (Eucoelosis) biloba (Linnaeus, 1767) Distribution: Mexico, Guatemala, Belize, El Salvador,

Honduras, Nicaragua, Costa Rica, Panama, Colombia, Venezuela, Guyana, Suriname, French Guiana, Brazil, Peru, Bolivia, Chile, and Paraguay (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (^{Ratcliffe & Cave, 2006: 245}); La Libertad: Teotepeque (^{Ratcliffe & Cave, 2006: 245}) (Fig. 24).

Years of collection: 1998-1999.

Months of collection: April and August.

Topographic zone: coastal mountains, coastal plain.

Elevation range: 520-780 m.

Comments: only 2 specimens from El Salvador were reported by ^{Ratcliffe and Cave (2006)}.

Enema Hope, 1837

Enema endymion Chevrolat, 1843

Distribution: Mexico, Belize, El Salvador, Honduras, Nicaragua, Costa Rica, Panama, Colombia, Venezuela, French Guiana, Brazil, and Bolivia (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (^{Ratcliffe & Cave, 2006: 248}); Santa Ana: El Congo (^{Ratcliffe & Cave, 2006: 249}); Parque Nacional Montecristo (Majaditas, San José Ingenio) (^{Serrano-Chicas, 2019: 37, 38}); Planes del Ranchador (^{Ratcliffe & Cave, 2006: 249}); Santa Ana (^{Ratcliffe & Cave, 2006: 248}); La Libertad: San Andrés (^{Ratcliffe & Cave, 2006: 249}); Santa Tecla (^{Berry & Salazar-Vaquero, 1957: 44}; ^{Ratcliffe & Cave, 2006: 249}); Volcán de San Salvador (^{Ratcliffe & Cave, 2006: 249}); San Salvador: Aguilares (^{Ratcliffe & Cave, 2006: 249}); Guazapa (^{Ratcliffe & Cave, 2006: 249}); San Salvador (^{Ratcliffe & Cave, 2006: 249}); Cuscatlán: San José Guayabal (^{Ratcliffe & Cave, 2006: 248}) (Fig. 24).

Years of collection: 1964, 1975-1979, 1983, 1988, 1997-2002, 2019.

Months of collection: All year, except October.

Topographic zone: northern mountains, interior valley, coastal mountains.

Elevation range: 300-1,845 m.

Heterogomphus Burmeister, 1847

Heterogomphus (Heterogomphus) chevrolatii Burmeister, 1847

Distribution: Mexico, Guatemala, El Salvador, Honduras, Nicaragua, Costa Rica, Panama, Colombia, Ecuador, and Brazil (^{Schoolmeesters, 2022}).

Locality records: Santa Ana: Parque Nacional Montecristo (Los Planes) (^{Ratcliffe & Cave, 2006: 255}); Chalatenango: Cerro El Pital (MUHNES); San Salvador: San Salvador (^{Ratcliffe & Cave, 2006: 255}); San Miguel: Moncagua (^{Ratcliffe & Cave, 2006: 255}); Morazán: Cerro Cacahuatique (^{Ratcliffe & Cave, 2006: 255}); Yamabal (^{Ratcliffe & Cave, 2006: 255}) (Fig. 24).

Years of collection: 1978, 1979, 1997-2002.

Months of collection: All year, except October.

Topographic zone: northern mountains, interior valley.

Elevation range: 245-2,660 m.

Heterogomphus (Daemonoplus) flohri (Kolbe, 1906) Distribution: Mexico, Belize, El Salvador, Guatemala, Honduras, and Nicaragua (^{Ratcliffe et al., 2013}). Locality records: Morazán: Cerro Cacahuatique (^{Ratcliffe & Cave, 2006: 258}) (Fig. 24).

Years of collection: 1979.

Months of collection: July.

Topographic zone: northern mountains.

Elevation range: 1,400 m.

Comments: only 2 specimens from El Salvador were reported by ^{Ratcliffe and Cave (2006)}, who misidentified them as Heterogomphus mniszechi (Thomson).

Irazua Ratcliffe, 2003

Irazua dilicra Ratcliffe, 2003

Distribution: El Salvador and Costa Rica (^{Schoolmeesters, 2022}).

Locality records: La Libertad: Santa Tecla (^{Ratcliffe & Cave, 2006: 258}) (Fig. 24). Years of collection: no data. Months of collection: May. Topographic zone: coastal mountains. Elevation range: 900 m.

Podischnus Burmeister, 1847 Podischnus agenor (Olivier, 1789)

Distribution: Mexico, Guatemala, Belize, El Salvador, Honduras, Nicaragua, Costa Rica, Panama, Colombia, Venezuela, Suriname, French Guiana, Ecuador, Brazil, and Peru (^{Schoolmeesters, 2022}).

Locality records: Santa Ana: Chalchuapa (^{Ratcliffe & Cave, 2006: 269}); La Libertad: Los Chorros (^{Ratcliffe & Cave, 2006: 269}); Quezaltepeque (^{Ratcliffe & Cave, 2006: 269}); San Andrés (^{Berry & Salazar-Vaquero, 1957: 47}; ^{Ratcliffe & Cave, 2006: 269}); San Juan Opico (^{Ratcliffe & Cave, 2006: 269}); Santa Tecla (^{Berry & Salazar-Vaquero, 1957: 47}; ^{Ratcliffe & Cave, 2006: 269}); San Salvador: Aguilares (^{Ratcliffe & Cave, 2006: 269}) (Fig. 24).

Years of collection: 1961, 1969, 1977-1978, 1986, 1997-2001.

Months of collection: April-September.

Topographic zone: interior valley, coastal mountains.

Elevation range: 300-900 m.

Comments:^{Berry and Salazar-Vaquero (1957)}, and ^{Berry (1959a)} stated that the species causes little damage to corn and sugarcane in El Salvador.

Strategus Hope, 1837

Strategus aloeus (Linnaeus, 1758)

Distribution: USA, Mexico, Guatemala, Belize, El Salvador, Honduras, Nicaragua, Costa Rica, Panama, Colombia, Venezuela, Trinidad & Tobago, Guyana, Suriname, French Guiana, Ecuador, Brazil, Peru, Bolivia, and Grenada (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Guaymango (^{Ratcliffe & Cave, 2006: 274}); Parque Nacional El Imposible (San Benito) (^{Ratcliffe & Cave, 2006: 274}; ^{Pablo-Cea, 2021: 22}; ^{Pablo-Cea, Deloya, MacGregor-Fors, Navarrete-Heredia, & Espinosa-de los Monteros, 2022: 379}); Santa Ana: Parque Nacional Montecristo (Bosque Nebuloso); (Los Planes) (^{Ratcliffe & Cave, 2006: 274}); Santa Ana (^{Ratcliffe & Cave, 2006: 274}); Sonsonate: Sonsonate (^{Ratcliffe & Cave, 2006: 274}); Chalatenango: La Montañona (^{Ratcliffe & Cave, 2006: 274}); La Libertad: El Barillo (MUHNES); La Libertad (^{Ratcliffe & Cave, 2006: 274}); Los Chorros (^{Ratcliffe & Cave, 2006: 274}); Parque El Bicentenario (^{Pablo-Cea, 2021: 22}; ^{Pablo-Cea, Deloya, MacGregor-Fors, & Navarrete-Heredia, 2022: 277}; ^{Pablo-Cea, Deloya, MacGregor-Fors, Navarrete-Heredia et al., 2022: 379}); San Andrés (^{Ratcliffe & Cave, 2006: 274}); Santa Tecla (^{Berry & Salazar-Vaquero, 1957: 47}; ^{Ratcliffe & Cave, 2006: 274}); Sitio del Niño (^{Ratcliffe & Cave, 2006: 274}); San Salvador: Aguilares (^{Ratcliffe & Cave, 2006: 274}); Cerro San Jacinto (^{Ratcliffe & Cave, 2006: 274}); Ilopango (^{Ratcliffe & Cave, 2006: 274}); Parque Saburo Hirao (MUHNES); Puerta del Diablo (^{Ratcliffe & Cave, 2006: 274}); San Salvador (^{Berry & Salazar-Vaquero, 1957: 47}; ^{Ratcliffe & Cave, 2006: 274}); Santo Tomás (^{Ratcliffe & Cave, 2006: 274}); Zacamil (^{Ratcliffe & Cave, 2006: 274}); Cuscatlán: Cojutepeque (^{Ratcliffe & Cave, 2006: 274}); El Rosario (^{Berry & Salazar-Vaquero, 1957: 47}); San José Guayabal (^{Ratcliffe & Cave, 2006: 274}); San Pedro Perulapán (^{Ratcliffe & Cave, 2006: 274}); La Paz: Ichanmichen (^{Ratcliffe & Cave, 2006: 274}); San Juan Talpa (^{Ratcliffe & Cave, 2006: 274}); Cabañas: Sensuntepeque (^{Ratcliffe & Cave, 2006: 274}); San Vicente: San Vicente (^{Ratcliffe & Cave, 2006: 274}); Santa Cruz Porrillo (^{Berry & Salazar-Vaquero, 1957: 47}); Usulután: Usulután (^{Ratcliffe & Cave, 2006: 274}); Morazán: Arambala (Río Sapo) (^{Ratcliffe & Cave, 2006: 274}) (Fig. 24).

Years of collection: 1958, 1964, 1972-1991, 1997-2002, 2019.

Months of collection: All year, except October.

Topographic zone: northern mountain, interior valley, coastal mountains, coastal plain.

Elevation range: 40-2,360 m.

Comments: reported as Strategus julianus Burmeister by ^{Berry and Salazar-Vaquero (1957)}.

Strategus hipposiderus Ratcliffe, 1976

Distribution: Mexico, Guatemala, El Salvador, Honduras, Nicaragua, Costa Rica, Panama, Colombia, and Brazil (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (^{Ratcliffe & Cave, 2006: 279}); Santa Ana: Parque Nacional Montecristo (Majaditas) (^{Serrano-Chicas, 2019: 37, 38}) (Fig. 24).

Years of collection: 1999, 2019.

Months of collection: April and August.

Topographic zone: northern mountains, coastal mountains.

Elevation range: 780-1,070 m.

Comments:^{Ratcliffe and Cave (2006)} reported only 1 specimen from El Salvador. ^{Serrano-Chicas (2019)} captured only 1 specimen, too.

Strategus jugurtha Burmeister, 1847

Distribution: Mexico, Belize, El Salvador, Honduras,

Nicaragua, Costa Rica, Panama, Colombia, Ecuador, and

Peru (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Ahuachapán (^{Ratcliffe & Cave, 2006: 282}); Parque Nacional El Imposible (San Benito) (^{Ratcliffe & Cave, 2006: 282}); Santa Ana: Parque Nacional Montecristo [probably San José Ingenio] (^{Ratcliffe & Cave, 2006: 282}; ^{Pablo-Cea, 2021: 22}; ^{Pablo-Cea, Deloya, MacGregor-Fors, Navarrete-Heredia, & Espinosa-de los Monteros, 2022: 379}); La Libertad: El Barillo (^{Ratcliffe & Cave, 2006: 282}); La Libertad (^{Ratcliffe & Cave, 2006: 282}); Parque El Bicentenario (^{Pablo-Cea, 2021: 22}; ^{Pablo-Cea, Deloya, MacGregor-Fors, & Navarrete-Heredia, 2022: 277}; ^{Pablo-Cea, Deloya, MacGregor-Fors, Navarrete-Heredia et al., 2022: 379}); San Andrés (^{Ratcliffe & Cave, 2006: 282}); Santa Tecla (^{Ratcliffe & Cave, 2006: 282}); Sitio del Niño (^{Ratcliffe & Cave, 2006: 282}); Zapotitán (^{Ratcliffe & Cave, 2006: 282}); San Salvador: Aguilares (^{Ratcliffe & Cave, 2006: 282}); San Salvador (^{Berry & Salazar-Vaquero, 1957: 47}; ^{Ratcliffe & Cave, 2006: 282}); Santo Tomás (^{Ratcliffe & Cave, 2006: 282}); Cuscatlán: Cojutepeque (^{Ratcliffe & Cave, 2006: 282}); Suchitoto (^{Ratcliffe & Cave, 2006: 282}); Cabañas: Cinquera (^{Ratcliffe & Cave, 2006: 282}); San Vicente: Volcán de San Vicente-Chinchontepec (^{Ratcliffe & Cave, 2006: 282}); Usulután: California (^{Ratcliffe & Cave, 2006: 282}); Estanzuelas (^{Ratcliffe & Cave, 2006: 282}); Usulután (^{Ratcliffe & Cave, 2006: 282}); Morazán: Arambala (Río Sapo) (^{Ratcliffe & Cave, 2006: 282}); La Unión: La Unión (^{Ratcliffe & Cave, 2006: 282}); Volcán de Conchagua (^{Ratcliffe & Cave, 2006: 282}) (Fig. 24). Years of collection: 1964, 1971-1993, 1997-2001, 2019. Months of collection: March-September. Topographic zone: northern mountain, interior valley, coastal mountains, coastal plain. Elevation range: 30-2,000 m.

Xyloryctes Hope, 1837

Xyloryctes ensifer Bates, 1888

Distribution: Mexico, Guatemala, El Salvador, and Honduras (^{Schoolmeesters, 2022}).

Locality records: Santa Ana: Parque Nacional Montecristo [probably Los Planes] (^{Ratcliffe & Cave, 2006: 288}); Chalatenango: Cerro El Pital (MUHNES); San Salvador: San Salvador (^{Ratcliffe & Cave, 2006: 288}) (Fig. 24).

Years of collection: 1975, 1978, 1979.

Months of collection: March and May.

Topographic zone: northern mountains, interior valley.

Elevation range: 685-2,660 m.

Comments:^{Ratcliffe and Cave (2006)} reported only 6 specimens collected in El Salvador.

Xyloryctes lobicollis Bates, 1888

Distribution: Mexico, Guatemala, El Salvador, Honduras, Nicaragua, Costa Rica, and Panama (^{School-meesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (^{Ratcliffe & Cave, 2006: 290}); Santa Ana; Parque Nacional Los Andes (^{Ratcliffe & Cave, 2006: 290}); Parque Nacional Montecristo (Los Planes, San José Ingenio) (^{Ratcliffe & Cave, 2006: 290}); La Libertad: San Andrés (^{Ratcliffe & Cave, 2006: 290}); Santa Tecla (^{Ratcliffe & Cave, 2006: 290}); San Salvador: Cuscatancingo (^{Ratcliffe & Cave, 2006: 290}); San Salvador (^{Berry & Salazar-Vaquero, 1957: 47}; ^{Ratcliffe & Cave, 2006: 290}); Cabañas: Cinquera (^{Ratcliffe & Cave, 2006: 290}); Sensuntepeque (^{Ratcliffe & Cave, 2006: 290}); La Unión: Volcán de Conchagua (MUHNES) (Fig. 25).

Years of collection: 1973-1979, 1985, 1997-2002.

Months of collection: March-November.

Topographic zone: northern mountains, interior valley, coastal mountains.

Elevation range: 385-1,890 m.

Tribe Pentodontini Mulsant, 1842 Euetheola Bates, 1888

Euetheola bidentata (Burmeister, 1847)

Distribution: Mexico, Guatemala, El Salvador, Honduras, Nicaragua, Costa Rica, Panama, Colombia, Venezuela, Trinidad & Tobago, Guyana, Suriname, French Guiana, and Brazil (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Atiquizaya (^{Ratcliffe & Cave, 2006: 197}); Santa Ana: Candelaria de la Frontera (^{Ratcliffe & Cave, 2006: 197}); La Libertad: Santa Tecla (^{Berry & Salazar-Vaquero, 1957: 45}); San Salvador: San Salvador (^{Ratcliffe & Cave, 2006: 197}); Cabañas: Cinquera (^{Ratcliffe & Cave, 2006: 197}); San Vicente: Santa Cruz Porrillo (^{Ratcliffe & Cave, 2006: 197}); Santo Domingo (FSCA); Usulután: Nancuchiname (^{Ratcliffe & Cave, 2006: 197}) (Fig. 25).

Years of collection: 1958-1969, 1978-1979, 1997-2002.

Months of collection: All year, except February and April.

Topographic zone: interior valley, coastal mountains, coastal plain.

Elevation range: 15-900 m.

Comments:^{Berry and Salazar-Vaquero (1957)} mentioned that adults feed on the leaves and stems of corn plants but cause little damage to the crop.

Euetheola humilis (Burmeister, 1847)

Distribution: Mexico, El Salvador, Honduras, Nicaragua, Costa Rica, Panama, Venezuela, Guyana, Suriname, French Guiana, Brazil, Bolivia, Paraguay, Argentina, and Uruguay (^{Schoolmeesters, 2022}).

Locality records: San Salvador: San Salvador (CMNC) (Fig. 25).

Years of collection: 1959.

Months of collection: July.

Topographic zone: interior valley.

Elevation range: 685 m.

Comments: new country record.

Ligyrus Casey, 1915

Ligyrus (Ligyrodes) sallaei Bates, 1888

Distribution: USA, Mexico, Guatemala, El Salvador, Honduras, Nicaragua, and Costa Rica (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (^{López-García & Deloya, 2022: 33}); (San Benito) (^{Ratcliffe & Cave, 2006: 238}; ^{Pablo-Cea, 2021: 22}; ^{Pablo-Cea, Deloya, MacGregor-Fors, Navarrete-Heredia, & Espinosa-de los Monteros, 2022: 379}); Santa Ana: Parque Nacional Los Andes (^{Ratcliffe & Cave, 2006: 238}); Parque Nacional Montecristo (Los Planes) (^{Ratcliffe & Cave, 2006: 238}); Chalatenango: La Montañona (^{Ratcliffe & Cave, 2006: 238}); La Libertad: San Andrés (^{Ratcliffe & Cave, 2006: 238}); San Salvador: San Salvador (^{Ratcliffe & Cave, 2006: 238}); Cabañas: Cinquera (^{Ratcliffe & Cave, 2006: 238}; Tejutepeque (El Tamagás) (MUHNES); Morazán: Cerro Cacahuatique (^{Ratcliffe & Cave, 2006: 238}); Yamabal (MUHNES); La Unión: Isla Martín Pérez (MUHNES); no specific department: no specific locality (^{Ratcliffe & Morón, 1997: 74}) (Fig. 25).

Years of collection: 1963, 1977-1978, 1994, 1997-2002.

Months of collection: March-November.

Topographic zone: northern mountain, interior valley, coastal mountains, coastal plain.

Elevation range: 40-1,890 m.

Comments: reported as Tomarus sallaei by ^{Ratcliffe and Cave (2006)}, ^{Pablo-Cea (2021)}, and ^{Pablo-Cea et al. (2022)}.

Ligyrus (Ligyrus) allonasutus^{López-García & Deloya, 2022}

Distribution: USA, Mexico, Guatemala, El Salvador, Honduras, Nicaragua, Costa Rica, Panama, and Jamaica (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Atiquizaya (Izcaquillo) (^{Ratcliffe & Cave, 2006: 237}); Guaymango (^{Ratcliffe & Cave, 2006: 237}); Parque Nacional El Imposible (San Benito) (^{Ratcliffe & Cave, 2006: 237}); Santa Ana: Parque Nacional Montecristo (Los Planes) (^{López-García & Deloya, 2022: 38}); (Majaditas) (^{Serrano-Chicas, 2019: 37, 38}); (San José Ingenio) (^{Ratcliffe & Cave, 2006: 237}; Sonsonate: Sonsonate (^{Ratcliffe & Cave, 2006: 237}); La Libertad: El Barillo (^{Ratcliffe & Cave, 2006: 237}); El Playón (^{López-García & Deloya, 2022: 38}); La Libertad (^{Ratcliffe & Cave, 2006: 237}); Parque Nacional Walter Thilo Deininger (^{Ratcliffe & Cave, 2006: 237}); San Andrés (^{Berry & Salazar-Vaquero, 1957: 45}; ^{Ratcliffe & Cave, 2006: 237}); San Diego (^{López-García & Deloya, 2022: 38}); Santa Tecla (^{Ratcliffe & Cave, 2006: 237}); Zapotitán (^{Ratcliffe & Cave, 2006: 237}); Zaragoza (^{Ratcliffe & Cave, 2006: 237}); San Salvador: Parque Saburo Hirao (^{López-García & Deloya, 2022: 38}); San Salvador (^{Ratcliffe & Cave, 2006: 237}); Cuscatlán: Cojutepeque (^{Ratcliffe & Cave, 2006: 237}); San Pedro Perulapán (^{Ratcliffe & Cave, 2006: 237}); La Paz: San Luis Talpa (^{López-García & Deloya, 2022: 38}); Zacatecoluca (^{Ratcliffe & Cave, 2006: 237}); Cabañas: Cinquera (^{Ratcliffe & Cave, 2006: 237}); Ilobasco (^{Ratcliffe & Cave, 2006: 237}); San Miguel: Laguna El Jocotal (^{López-García & Deloya, 2022: 38}); Morazán: Arambala (Río Sapo) (^{Ratcliffe & Cave, 2006: 237}); La Unión: La Unión (^{Ratcliffe & Cave, 2006: 237}) (Fig. 25).

Years of collection: 1952, 1964, 1972-1979, 1983-1990, 1997-2002, 2019.

Months of collection: All year.

Topographic zone: northern mountain, interior valley, coastal mountains, coastal plain.

Elevation range: 15-1,890 m.

Comments: reported as Tomarus nassutus [sic] by ^{Serrano-Chicas (2019)} and as Tomarus nasutus by ^{Berry and Salazar-Vaquero (1957)}, ^{Berry (1959a)}, and ^{Ratcliffe and Cave (2006)}. ^{Berry and Salazar-Vaquero (1957)}, and ^{Berry (1959a)} mentioned that the species is harmful to cultivated lettuce in El Salvador. ^{López-García and Deloya (2022)} examined specimens from La Paz: Caprex, a locality unknown to us.

Ligyrus (Ligyrus) fossor (Latreille, 1813)

Distribution: El Salvador, Panama, Colombia, Venezuela, Ecuador, Brazil, Bolivia, Bahamas, Cuba, Jamaica, Puerto Rico, Antigua & Barbuda, and Curaçao (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (MUHNES); (San Francisco Menéndez) (^{Ratcliffe & Cave, 2006: 228}); La Unión: Isla Martín Pérez (^{Ratcliffe & Cave, 2006: 228}) (Fig. 25).

Years of collection: 2001.

Months of collection: July and September.

Topographic zone: coastal mountains, coastal plain.

Elevation range: 40-780 m.

Comments: reported as Tomarus fossor in ^{Ratcliffe and Cave (2006)}, who reported only 3 specimens from El Salvador.

Orizabus Fairmaire, 1878 Orizabus clunalis (LeConte, 1856)

Distribution: USA, Mexico, Guatemala, El Salvador, and Honduras (^{Schoolmeesters, 2022}).

Locality records: Santa Ana: Parque Nacional Montecristo (Bosque Nebuloso) (^{Ratcliffe & Cave, 2006: 208}; ^{Serrano-Chicas, 2019: 37, 38}) (Fig. 25).

Years of collection: 2019.

Months of collection: March and April.

Topographic zone: northern mountains.

Elevation range: 2,360 m.

Comments: reported as Orizabus clunaris [sic] by ^{Serrano-Chicas (2019)}. Only 1 specimen from El Salvador was reported by ^{Ratcliffe and Cave (2006)}, with no specified year and month of collection. ^{Serrano-Chicas (2019)} collected 4 specimens in the cloud forest of Parque Nacional Montecristo.

Orizabus teamscaraborum^{Ratcliffe & Cave, 2006} Distribution: El Salvador and Honduras (^{School-meesters, 2022}).

Locality records: Santa Ana: Parque Nacional Montecristo (Bosque Nebuloso) (^{Ratcliffe & Cave, 2006: 215}); Chalatenango: Cerro El Pital (^{Ratcliffe & Cave, 2006: 215}) (Fig. 25).

Years of collection: 1960, 1971, 1997-2002.

Months of collection: February, April-July, and October.

Topographic zone: northern mountains.

Elevation range: 2,360-2,660 m.

Proculigyrus López-García & Deloya, 2022 Proculigyrus cicatricosus (Prell, 1937)

Distribution: Mexico, Guatemala, El Salvador, Honduras, Costa Rica, and Colombia (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (^{Ratcliffe & Cave, 2006: 225}); San Miguelito (^{López-García & Deloya, 2022: 22}); Santa Ana: Parque Nacional Montecristo (^{López-García & Deloya, 2022: 22}) (Bosque Nebuloso, Cerro Miramundo) (^{López-García & Deloya, 2022: 22}); [probably San José Ingenio] (^{Ratcliffe & Cave, 2006: 225}); Metapán (^{López-García & Deloya, 2022: 22}); La Libertad: El Barillo (^{Ratcliffe & Cave, 2006: 225}); San Andrés (^{Ratcliffe & Cave, 2006: 225}); Santa Tecla (^{Ratcliffe & Cave, 2006: 225}); San Salvador: San Salvador (^{López-García & Deloya, 2022: 22}) (Fig. 25).

Years of collection: 1952, 1975-1980, 1997-2002.

Months of collection: February, March, May-July, and September-December.

Topographic zone: northern mountains, interior valley, coastal mountains.

Elevation range: 475-2,360 m.

Comments: reported as Tomarus cicatricosus by ^{Ratcliffe and Cave (2006)}.

Tomarus Erichson, 1847

Tomarus bituberculatus (Palisot de Beauvois, 1811) Distribution: Mexico, Guatemala, El Salvador, Honduras, Nicaragua, Costa Rica, Panama, Colombia, Venezuela, Trinidad & Tobago, Ecuador, Brazil, Peru, and Bolivia (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (La Fincona, San Benito) (MUHNES); San Miguelito (MUHNES); Santa Ana: Metapán (MUHNES); Parque Nacional Montecristo (Los Planes, San José Ingenio) (MUHNES); San Salvador: Parque Saburo Hirao (MUHNES); San Salvador (^{Ratcliffe & Cave, 2006: 221}); (Scandia) (MUHNES); Cuscatlán: Área Natural Protegida Colima (Cerrón Grande) (MUHNES); San Vicente: no specific locality (MUHNES); La Unión: no specific locality (MUHNES) (Fig. 25).

Years of collection: 1964, 1972-1977.

Months of collection: May, June, August, and November.

Topographic zone: northern mountains, interior valley, coastal mountains.

Elevation range: 305-1,890 m.

Tomarus discrepans (Escalona & Joly, 2006) Distribution: Mexico, Guatemala, El Salvador,

Honduras, Nicaragua, Costa Rica, Panama, and Venezuela (^{López-García & Deloya 2022}).

Locality records: Santa Ana: Santa Ana (^{Ratcliffe & Cave, 2006: 229}); La Libertad: San Andrés (^{Ratcliffe & Cave, 2006: 228}); Santa Tecla (^{Ratcliffe & Cave, 2006: 228}); San Salvador: San Salvador (^{Ratcliffe & Cave, 2006: 228}); Usulután: Usulután (^{Ratcliffe & Cave, 2006: 228}) (Fig. 26).

Years of collection: 1952, 1975-1979.

Months of collection: April, September, October, and December.

Topographic zone: interior valley, coastal mountains, coastal plain.

Elevation range: 80-900 m.

Comments: reported as Tomarus gyas Erichson by ^{Ratcliffe and Cave (2006)}. Although the species is common in Honduras, ^{Ratcliffe and Cave (2006)} reported only 6 specimens from El Salvador.

Tomarus laevicollis (Bates, 1888)

Distribution: Mexico, El Salvador, Honduras, Nicaragua, and Costa Rica (^{Schoolmeesters, 2022}).

Locality records: La Paz: Playa Costa del Sol (MUHNES); San Miguel: Laguna El Jocotal (^{Ratcliffe & Cave, 2006: 232}) (Fig. 26).

Years of collection: 2001.

Months of collection: May.

Topographic zone: coastal plain.

Elevation range: 5-25 m.

Comments:^{Ratcliffe and Cave (2006)} reported only 1 specimen from El Salvador.

Tribe Phileurini Burmeister, 1847

Goniophileurus Kolbe, 1910

Goniophileurus femoratus (Burmeister, 1847)

Distribution: Mexico, El Salvador, Honduras, Costa Rica, Panama, Colombia, Venezuela, French Guiana, Ecuador, and Brazil (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (^{Ratcliffe & Cave, 2006: 308}); Morazán: Arambala (Río Sapo) (^{Ratcliffe & Cave, 2006: 308}) (Fig. 26).

Years of collection: 1999, 2001.

Months of collection: May.

Topographic zone: northern mountains, coastal mountains.

Elevation range: 665-780 m.

Comments:^{Ratcliffe and Cave (2006)} reported only 2 specimens from El Salvador.

Hemiphileurus Kolbe, 1910 Hemiphileurus cylindroides (Bates, 1888)

Distribution: Mexico, Guatemala, Belize, El Salvador, Nicaragua, Costa Rica, and Panama (^{Schoolmeesters, 2022}).

Locality records: Cuscatlán: Suchitoto (^{Ratcliffe & Cave, 2006: 314}) (Fig. 26).

Years of collection: 1979.

Months of collection: September.

Topographic zone: interior valley.

Elevation range: 385 m.

Comments:^{Ratcliffe and Cave (2006)} reported only 1 specimen from El Salvador.

Hemiphileurus euniceae Ratcliffe & Cave, 2006 Distribution: Guatemala and El Salvador

(^{Schoolmeesters, 2022}).

Locality records: San Salvador: Soyapango (^{Ratcliffe & Cave, 2006: 317}) (Fig. 26). Years of collection: 1999. Months of collection: June. Topographic zone: interior valley. Elevation range: 610 m.

Comments: only 1 specimen from El Salvador is known (^{Ratcliffe & Cave, 2006}). Hemiphileurus laevicauda (Bates, 1888)

Distribution: Mexico, Guatemala, El Salvador, Honduras, Nicaragua, Costa Rica, and Panama (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (MUHNES); Santa Ana: Santa Ana (^{Ratcliffe & Cave, 2006: 318}); Chalatenango: San Ignacio (^{Ratcliffe & Cave, 2006: 318}); San Salvador: Parque Saburo Hirao (MUHNES); San Vicente: Apastepeque (Poza Azul) (MUHNES) (Fig. 26).

Years of collection: 1977-1979, 1998.

Months of collection: March, May, and June.

Topographic zone: northern mountains, interior valley, coastal mountains.

Elevation range: 500-1,060 m.

Homophileurus Kolbe, 1910 Homophileurus luedeckei Kolbe, 1910

Distribution: Mexico, Guatemala, and El Salvador (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (^{Ratcliffe & Cave, 2006: 327}); San Miguelito (near the ecotone of Parque Nacional El Imposible) (E. Sandoval, personal communication) (Fig. 26).

Years of collection: 1979, 2013.

Months of collection: June and November.

Topographic zone: coastal mountains.

Elevation range: 725-800 m.

Comments:^{Ratcliffe and Cave (2006)} reported only a single specimen from El Salvador. Eliberto Sandoval collected a specimen in November 2013 (E. Sandoval, personal communication).

Homophileurus quadrituberculatus (Palisot de Beauvois, 1805)

Distribution: Mexico, Guatemala, Belize, El Salvador, Nicaragua, Costa Rica, Panama, Colombia, Venezuela, Guyana, Suriname, French Guiana, Ecuador, Brazil, Peru, Bolivia, Haiti, Dominican Republic, Puerto Rico, Dominica, and Grenada (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (^{Ratcliffe & Cave, 2006: 328}); Santa Ana: Parque Nacional Montecristo (Los Planes) (MUHNES) (Fig. 26).

Years of collection: no data.

Months of collection: no data.

Topographic zone: northern mountains, coastal mountains.

Elevation range: 780-1,890 m.

Comments:^{Ratcliffe and Cave (2006)} reported only a single specimen from El Salvador, with no specified year and month of collection.

Homophileurus tricuspis Prell, 1914

Distribution: Mexico, Guatemala, Belize, El Salvador, Honduras, Nicaragua, Costa Rica, Panama, Colombia, Venezuela, and Brazil (^{Schoolmeesters, 2022}).

Locality records: Cabañas: Cinquera (^{Ratcliffe & Cave, 2006: 331}) (Fig. 26).

Years of collection: no data.

Months of collection: June.

Topographic zone: interior valley.

Elevation range: 385 m.

Comments:^{Ratcliffe and Cave (2006)} reported only 1 specimen from El Salvador, with no specified year of collection.

Phileurus Latreille, 1807

Phileurus didymus (Linnaeus, 1758)

Distribution: Mexico, Guatemala, El Salvador, Honduras, Nicaragua, Costa Rica, Panama, Colombia, Venezuela, Trinidad & Tobago, Guyana, Suriname, French Guiana, Ecuador, Brazil, Peru, Bolivia, Paraguay, Puerto Rico, Guadeloupe, Dominica, Martinique, St. Lucia, St. Vincent & the Grenadines, and Grenada (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Apaneca (^{Ratcliffe & Cave, 2006: 337}); Parque Nacional El Imposible (San Benito) (^{Ratcliffe & Cave, 2006: 337}; ^{Pablo-Cea, 2021: 22}; ^{Pablo-Cea, Deloya, MacGregor-Fors, Navarrete-Heredia, & Espinosa-de los Monteros, 2022: 379}); Santa Ana: El Congo (^{Ratcliffe & Cave, 2006: 337}); Parque Nacional Montecristo (Majaditas) (^{Serrano-Chicas, 2019: 37, 38}); (San José Ingenio) (^{Ratcliffe & Cave, 2006: 337}; ^{Serrano-Chicas, 2019: 37, 38}); Sonsonate: Sonsonate (^{Ratcliffe & Cave, 2006: 337}); Chalatenango: Chalatenango (^{Ratcliffe & Cave, 2006: 337}); La Libertad: Los Chorros (^{Ratcliffe & Cave, 2006: 337}); San Andrés (^{Ratcliffe & Cave, 2006: 337}); Santa Tecla (^{Ratcliffe & Cave, 2006: 337}); Zapotitán (^{Ratcliffe & Cave, 2006: 337}); San Salvador: San Salvador (^{Ratcliffe & Cave, 2006: 337}); Cabañas: Cinquera (^{Ratcliffe & Cave, 2006: 337}) (Fig. 26).

Years of collection: 1959, 1969, 1974, 1987, 1994, 1997-2002, 2019.

Months of collection: All year.

Topographic zone: northern mountains, interior valley, coastal mountains, coastal plain.

Elevation range: 225-1,475 m.

Phileurus valgus (Olivier, 1789)

Distribution: USA, Mexico, Guatemala, Belize, El Salvador, Honduras, Nicaragua, Costa Rica, Panama, Colombia, Venezuela, Trinidad & Tobago, Guyana, Suriname, French Guiana, Brazil, Peru, Bolivia, Paraguay, Argentina, Bahamas, Cuba, Cayman Islands, Jamaica, Dominican Republic, Puerto Rico, US Virgin Islands, St. Kitts & Nevis, Antigua & Barbuda, Montserrat, Guadeloupe, Dominica, Martinique, St. Lucia, St. Vincent & the Grenadines, Barbados, and Grenada (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (^{Ratcliffe & Cave, 2006: 342}; ^{Pablo-Cea, 2021: 22}; ^{Pablo-Cea, Deloya, MacGregor-Fors, Navarrete-Heredia, & Espinosa-de los Monteros, 2022: 379}); Santa Ana: Parque Nacional Montecristo (Los Planes) (^{Ratcliffe & Cave, 2006: 342}); (San José Ingenio) (^{Serrano-Chicas, 2019: 37, 38}); Santa Ana (^{Ratcliffe & Cave, 2006: 342}); La Libertad; Parque Nacional Walther Thilo Deininger (^{Ratcliffe & Cave, 2006: 342}); San Salvador: Ilopango (^{Ratcliffe & Cave, 2006: 342}); San Salvador (^{Ratcliffe & Cave, 2006: 342}); San Miguel: Laguna El Jocotal (^{Ratcliffe & Cave, 2006: 342}); Morazán: Arambala (Río Sapo) (^{Ratcliffe & Cave, 2006: 342}) (Fig. 27).

Years of collection: 1977, 1988, 1997-2002, 2019.

Months of collection: All year.

Topographic zone: northern mountains, interior valley, coastal mountains, coastal plain.

Elevation range: 25-1890 m.

Comments: reported as Phileurus valgas [sic] by ^{Serrano-Chicas (2019)}.

Subfamily Cetoniinae Leach, 1815 Tribe Cetoniini Leach, 1815 Euphoria Burmeister, 1842 Euphoria avita Janson, 1881

Distribution: Mexico, Guatemala, El Salvador, and Honduras (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (^{Pablo-Cea, 2021: 22}; ^{Pablo-Cea, Deloya, MacGregor-Fors, Navarrete-Heredia, & Espinosa-de los Monteros, 2022: 379}); Santa Ana: Parque Nacional Cerro Verde (^{Orozco, 2012: 62}); Parque Nacional Los Andes (^{Orozco, 2012: 62}); Chalatenango: La Palma (^{Orozco, 2012: 62}) (Fig. 27).

Years of collection: 2019.

Months of collection: May and June.

Topographic zone: northern mountains, coastal mountains.

Elevation range: 780-2,020 m.

Comments: in Parque Nacional El Imposible, specimens were captured with aerial traps baited with fermented fruit (^{Pablo-Cea, 2021}).

Euphoria biguttata (Gory & Percheron, 1833) Distribution: USA, Mexico, Guatemala, El Salvador,

and Honduras (^{Schoolmeesters, 2022}).

Locality records: San Salvador: San Salvador (^{Orozco, 2012: 18}); Cuscatlán: El Rosario (^{Orozco, 2012: 18}) (Fig. 27).

Years of collection: no data.

Months of collection: no data.

Topographic zone: interior valley.

Elevation range: 685-720 m.

Euphoria canescens (Gory & Percheron, 1833) Distribution: USA, Mexico, Guatemala, Belize, El

Salvador, and Honduras (^{Schoolmeesters, 2022}). Locality records: Ahuachapán: Parque Nacional El Imposible [probably San Benito] (^{Orozco, 2012: 19});

Cuscatlán: El Rosario (^{Berry & Salazar-Vaquero, 1957: 45}) (Fig. 27).

Years of collection: 1979.

Months of collection: November.

Topographic zone: interior valley, coastal mountains.

Elevation range: 720-780 m.

Euphoria dimidiata (Gory & Percheron, 1833) Distribution: USA, Mexico, Guatemala, El Salvador,

and Honduras (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: 5 km NE of San Francisco Menéndez (^{Orozco, 2012: 21}); La Libertad: Santa Tecla (^{Orozco, 2012: 21}); San Salvador: San Salvador (TAMU); Cuscatlán: El Rosario (^{Berry & Salazar-Vaquero, 1957: 45}; ^{Orozco, 2012: 21}); San Miguel: La Ceiba (^{Orozco, 2012: 21)} (Fig. 27).

Years of collection: 1979.

Months of collection: November.

Topographic zone: interior valley, coastal mountains, coastal plain.

Elevation range: 170-900 m. Euphoria geminata (Chevrolat, 1834)

Distribution: Mexico, Guatemala, El Salvador, Honduras, and Nicaragua (^{Schoolmeesters, 2022}).

Locality records: Santa Ana: Candelaria [de la Frontera]

(^{Orozco, 2012: 77}); Parque Nacional Montecristo (Los Planes) (MUHNES); Sonsonate: Izalco (Los Guates Farm) (MUHNES); Sonsonate (^{Orozco, 2012: 77}); La Libertad: Santa Tecla (^{Orozco, 2012: 77}); San Salvador: Cerro San Jacinto (MUHNES); Cuscatlán: El Rosario (^{Berry & Salazar-Vaquero, 1957: 45}; ^{Orozco, 2012: 77}); La Paz: Mercedes La Ceiba (^{Orozco, 2012: 77}) (Fig. 27).

Years of collection: 1972, 1974-1976.

Months of collection: May and November.

Topographic zone: northern mountains, interior valley, coastal mountains, coastal plain.

Elevation range: 225-1,890 m. Euphoria iridescens (Schaum, 1841)

Distribution: Mexico, Guatemala, El Salvador, Honduras, Nicaragua, and Costa Rica (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (^{Pablo-Cea, 2021: 22}; ^{Pablo-Cea, Deloya, MacGregor-Fors, Navarrete-Heredia, & Espinosa-de los Monteros, 2022: 379}) (Fig. 27).

Years of collection: 2018, 2019.

Months of collection: February, and April-September.

Topographic zone: coastal mountains.

Elevation range: 780 m.

Comments: adults were captured in aerial traps baited with fermented fruit, and 1 adult was found in a pitfall trap baited with human dung (^{Pablo-Cea, 2021}).

Euphoria lesueuri (Gory & Percheron, 1833) Distribution: Mexico, Guatemala, Belize, El Salvador,

Honduras, and Nicaragua (^{Schoolmeesters, 2022}). Locality records: Santa Ana: Parque Nacional Montecristo (San José Ingenio) (^{Orozco, 2012: 55}); La Libertad: Santa Tecla (Orozco, 2012: 55) (Fig. 27).

Years of collection: 1979.

Months of collection: April.

Topographic zone: northern mountains, coastal mountains.

Elevation range: 810-900 m.

Euphoria pulchella (Gory & Percheron, 1833) Distribution: Mexico, Guatemala, Belize, El Salvador,

Honduras, Nicaragua, Costa Rica, and Venezuela (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (^{Pablo-Cea, 2021: 22}; ^{Pablo-Cea, Deloya, MacGregor-Fors, Navarrete-Heredia, & Espinosa-de los Monteros, 2022: 379}); San Salvador: San Salvador (^{Orozco, 2012: 91}); Cuscatlán: El Rosario (^{Berry & Salazar-Vaquero, 1957: 45}; ^{Orozco, 2012: 91}) (Fig. 27). Years of collection: 2019.

Months of collection: May and June.

Topographic zone: interior valley, coastal mountains.

Elevation range: 685-780 m.

Euphoria yucateca Bates, 1889

Distribution: Mexico, Guatemala, Belize, El Salvador, Honduras, Nicaragua, Costa Rica, and Venezuela (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (^{Pablo-Cea, 2021: 22}; ^{Pablo-Cea, Deloya, MacGregor-Fors, Navarrete-Heredia, & Espinosa-de los Monteros, 2022: 379}); La Libertad: Santa Tecla (^{Berry & Salazar-Vaquero, 1957: 45}; ^{Orozco, 2012: 41}); San Salvador: San Salvador (^{Orozco, 2012: 41}); Cuscatlán: El Rosario (^{Berry & Salazar-Vaquero, 1957: 45}; ^{Orozco, 2012: 41}); Cabañas: Cinquera (^{Orozco, 2012: 41}); San Miguel: San Jacinto (^{Orozco, 2012: 41}); San Miguel (^{Orozco, 2012: 41}) (Fig. 27).

Years of collection: 1979, 2019.

Months of collection: May, June, and September.

Topographic zone: interior valley, coastal mountains.

Elevation range: 135-900 m.

Comments: reported as Euphoria limatula Janson and E. yucateca by ^{Berry and Salazar-Vaquero (1957)} and as E. limatula by ^{Berry (1959b)} and ^{Solís (2004)}. Erroneously identified as E. leucigrapha [sic] by ^{Andrews et al. (1979)}. ^{Berry and Salazar-Vaquero (1957)} mentioned that adults feed on rosebush and “guineo enano” (Musa sp). In Parque Nacional El Imposible, specimens were captured in aerial traps baited with fermented fruit (^{Pablo-Cea, 2021}).

Tribe Gymnetini Kirby, 1827

Amithao Thomson, 1878

Amithao albopictus Neervoort van de Poll, 1886 Distribution: El Salvador, Nicaragua, Costa Rica, and Panama (^{Pablo-Cea & Alfaro, 2020}; ^{Schoolmeesters, 2022}).

Locality records: San Salvador: Ilopango (^{Pablo-Cea & Alfaro, 2020: 24}) (Fig. 28). Years of collection: 2010. Months of collection: May. Topographic zone: interior valley. Elevation range: 625 m.

Amithao anthracinus^{Ratcliffe, 2013}

Distribution: El Salvador and Panama (^{Pablo-Cea & Alfaro, 2020}; ^{Schoolmeesters, 2022}).

Locality records: San Salvador: San Salvador (Universidad de El Salvador) (^{Pablo-Cea & Alfaro, 2020: 24}) (Fig. 28).

Years of collection: 2019.

Months of collection: May.

Topographic zone: interior valley.

Elevation range: 695 m.

Argyripa Thomson 1878 Argyripa lansbergei (Sallé, 1857)

Distribution: México, El Salvador, Nicaragua, Colombia, Ecuador, and Brazil (^{Pablo-Cea & Alfaro, 2020}; ^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (^{Pablo-Cea, 2021: 22}; ^{Pablo-Cea, Deloya, MacGregor-Fors, Navarrete-Heredia, & Espinosa-de los Monteros, 2022: 379}) (Fig. 28).

Years of collection: 2019.

Months of collection: December.

Topographic zone: coastal mountains.

Elevation range: 780 m.

Cotinis Burmeister, 1842

Cotinis (Cotinis) mutabilis (Gory & Percheron, 1833) Distribution: USA, Mexico, Guatemala, Belize, El Salvador, Honduras, Nicaragua, and Costa Rica (^{Solís, 2004}; ^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible [probably San Benito] (RDCC); Santa Ana: Parque Nacional Montecristo (Los Planes) (MUHNES); Santa Ana (^{Berry & Salazar-Vaquero, 1957: 44}); Sonsonate: Izalco (Los Guates Farm) (MUHNES); La Libertad: Los Chorros (MUHNES); Parque El Bicentenario (^{Pablo-Cea, 2021}; ^{Pablo-Cea, Deloya, MacGregor-Fors, & Navarrete-Heredia, 2022: 277}; ^{Pablo-Cea, Deloya, MacGregor-Fors, Navarrete-Heredia et al., 2022: 379}); Parque Nacional Walter Thilo Deininger (MUHNES); San Salvador: Cerro San Jacinto (J. Pablo, personal observation); Delgado (RDCC); Mejicanos (A. Pablo, personal communication); Parque Saburo Hirao (MUHNES); San Salvador (RDCC; S. Martínez, personal communication) (Fig. 28).

Years of collection: 1975-1979, 2008-2009, 2018-2019.

Months of collection: April, and June-November.

Topographic zone: northern mountains, interior valley, coastal mountains, coastal plain.

Elevation range: 135-1,890 m.

Comments: widespread in El Salvador, with surprisingly few locality records in the literature. In El Salvador, the species is known by the common name “ron ron” (^{Berry & Salazar-Vaquero, 1957}). Children in rural zones used to play with tethered adults, making them flying pets (J. F. Pablo, personal communication). In Parque El Bicentenario, specimens were captured with aerial traps baited with fermented fruit (^{Pablo-Cea, 2021}). ^{Berry and Salazar-Vaquero (1957)} mentioned that the species is harmful to cultivated pineapple. ^{Berry (1959a)} stated that the species is commonly found feeding on fruits in cultivated citrus in El Salvador but is not harmful to the trees. One specimen was observed feeding on a peach fruit (^{Berry 1959a}).

Gymnetis MacLeay, 1819

Gymnetis bajula wollastoni Schaum, 1848

Distribution: Mexico, Guatemala, El Salvador, Honduras, Nicaragua, Costa Rica, and Panama (^{Ratcliffe, 2018}; ^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (^{Ratcliffe, 2018: 40}); La Paz: no specific locality (^{Ratcliffe, 2018: 40}) (Fig. 28).

Years of collection: no data.

Months of collection: no data.

Topographic zone: coastal mountains.

Elevation range: 780 m.

Gymnetis ramulosa Bates, 1869

Distribution: Belize, Guatemala, El Salvador, Honduras, Nicaragua, Costa Rica, Panama, Colombia, Brazil, and Venezuela (^{Solís, 2004}; ^{Ratcliffe, 2018}; ^{Pablo-Cea & Alfaro, 2020}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (^{Pablo-Cea, 2021: 22}; ^{Pablo-Cea, Deloya, MacGregor-Fors, Navarrete-Heredia, & Espinosa-de los Monteros, 2022: 379}); La Libertad: Área Natural Protegida San Andrés (J. Pablo, personal observation); Parque El Bicentenario (^{Pablo-Cea, 2021: 22}; ^{Pablo-Cea, Deloya, MacGregor-Fors, & Navarrete-Heredia, 2022: 277}; ^{Pablo-Cea, Deloya, MacGregor-Fors, Navarrete-Heredia et al., 2022: 379}) (Fig. 28).

Years of collection: 2017-2019.

Months of collection: February, and May-July.

Topographic zone: interior valley, coastal mountains.

Elevation range: 475-870 m.

Comments: reported as Gymnetis chevrolati Gory & Percheron by ^{Pablo-Cea & Alfaro (2020)}. The specimen captured in the surroundings of the Área Natural Protegida San Andrés was feeding on the fruit of a cashew tree (Anacardium occidentale L.) (J. Pablo, personal observation). In Parque Nacional El Imposible and Parque El Bicentenario, the specimens were captured in aerial traps baited with fruit (^{Pablo-Cea, 2021}).

Gymnetis stellata (Latreille, 1833)

Distribution: Mexico, Guatemala, Belize, El Salvador, Honduras, Nicaragua, Costa Rica, Panama, Colombia, Venezuela, Ecuador, Brazil, Peru, and Dominican Republic (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Barra de Santiago (^{Ratcliffe, 2018: 205}); San Salvador: San Salvador (^{Ratcliffe, 2018: 205}) (Fig. 28).

Years of collection: 1980.

Months of collection: January.

Topographic zone: interior valley, coastal plain.

Elevation range: 5-685 m.

Comments. Label data record the specimen from Barra de Santiago was “feeding on a cut portion of botoncillo (probably Conocarpus erectus L.)” (R. D. Cave, personal observation).

Hologymnetis Martínez, 1949

Hologymnetis cinerea (Gory & Percheron, 1833) Distribution: USA?, Mexico, Guatemala, El Salvador, Honduras, and Nicaragua (^{Delgado & Márquez, 2006}; ^{Maes & Ratcliffe, 2020}; ^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible [probably San Benito] (RDCC); La Libertad: Los Chorros (RDCC); no specific department: Cafetalera? (^{Ratcliffe & Deloya, 1992: 169}) (Fig. 28).

Years of collection: 1979.

Months of collection: September and November.

Topographic zone: coastal mountains.

Elevation range: 730-780 m. Hologymnetis margaritis (Bates, 1889)

Distribution: Mexico, Guatemala, and El Salvador (^{Ratcliffe & Deloya, 1992}; ^{Schoolmeesters, 2022}).

Locality records: La Libertad: Santa Tecla (^{Berry & Salazar-Vaquero, 1957: 45}) (Fig. 28).

Years of collection: no data.

Months of collection: no data.

Topographic zone: coastal mountains.

Elevation range: 900 m.

Comments: reported as Gymnetis margaritis by ^{Berry and Salazar-Vaquero (1957)}, and ^{Berry (1959a)}. ^{Berry and Salazar-Vaquero (1957)}, and ^{Berry (1959a)} mentioned that the species is common in the coffee plantations of El Salvador, but it is not harmful to the plants.

Hologymnetis vulcanorum^{Ratcliffe & Deloya, 1992}Distribution: Guatemala and El Salvador (^{Ratcliffe & Deloya, 1992}; ^{Schoolmeesters, 2022}).

Locality records: Santa Ana: Candelaria [de la Frontera] (^{Ratcliffe & Deloya, 1992: 179}); La Libertad: Santa Tecla (^{Ratcliffe & Deloya, 1992: 179}); Cuscatlán: El Rosario (^{Ratcliffe & Deloya, 1992: 179}) (Fig. 28).

Years of collection: 1951-1953.

Months of collection: August and September.

Topographic zone: interior valley, coastal mountain.

Elevation range: 720-900 m.

Hoplopyga Thomson, 1880 Hoplopyga liturata (Olivier, 1789)

Distribution: Mexico, Guatemala, Belize, El Salvador, Honduras, Nicaragua, Costa Rica, Panama, Colombia, Venezuela, Trinidad & Tobago, Guyana, Suriname, French Guiana, Ecuador, Brazil, Peru, Bolivia, Chile, Paraguay, and Argentina (^{Schoolmeesters, 2022}).

Locality records: Ahuachapán: Parque Nacional El Imposible (San Benito) (^{Pablo-Cea, 2021: 22}; ^{Pablo-Cea, Deloya, MacGregor-Fors, Navarrete-Heredia, & Espinosa-de los Monteros, 2022: 379}); La Libertad: Parque El Bicentenario (^{Pablo-Cea, 2021: 22}; ^{Pablo-Cea, Deloya, MacGregor-Fors, & Navarrete-Heredia, 2022: 277}; ^{Pablo-Cea, Deloya, MacGregor-Fors, Navarrete-Heredia et al., 2022: 379}); Santa Tecla (^{Berry & Salazar-Vaquero, 1957: 45}); San Salvador: Los Planes de Renderos (^{Shaughney & Ratcliffe, 2015: 608}); San Miguel: no specific locality (^{Shaughney & Ratcliffe, 2015: 608}) (Fig. 29).

Years of collection: 2019.

Months of collection: January, April-September, November, and December.

Topographic zone: coastal mountains.

Elevation range: 780-1,005 m.

Comments: reported as Hoplopyga literata [sic] by ^{Berry and Salazar-Vaquero (1957)}, and as Gymnatis [sic] liturata in ^{Andrews et al. (1979)}. The specimens from Parque Nacional El Imposible and Parque El Bicentenario were captured in aerial traps baited with fruit (^{Pablo-Cea, 2021}).

Tribe Trichiini Fleming, 1821 Trigonopeltastes Burmeister & Schaum, 1840 Trigonopeltastes archimedes Schaum in Burmeister & Schaum, 1841

Distribution: Mexico, Guatemala, El Salvador, and Costa Rica (^{Smith, 2016}; ^{Schoolmeesters, 2022}).

Locality records: La Unión: Volcán de Conchagua (^{Howden, 1968: 48}) (Fig. 29).

Years of collection: 1958.

Months of collection: May.

Topographic zone: coastal mountains.

Elevation range: 1,190 m.

Comments: it is likely that the year and month of collection for the specimen from La Unión is the same (May 1958) as that of Trigonopeltastes variabilis Howden, since both species are reported in the same paper (^{Howden, 1968}).

Trigonopeltastes frontalis Bates, 1889

Distribution: Mexico, Belize, Guatemala, El Salvador, Nicaragua, and Honduras (^{Smith, 2016}).

Locality records: Ahuachapán: Parque Nacional El Imposible (Cerro de Loma de Paja, Puerta Mayor [Sector San Francisco Menéndez]) (^{Cave, 1983: 152}) (Fig. 29).

Years of collection: 1979.

Months of collection: June.

Topographic zone: coastal plain.

Elevation range: 250 m.

Comments: adults were found on or flying around the flowers of Casearia aculeata Jacq. (^{Cave, 1983}). Trigonopeltastes geometricus Schaum in Burmeister & Schaum, 1841

Distribution: Mexico, Belize, Guatemala, El Salvador, Honduras, Nicaragua, Costa Rica, Panama, Venezuela, Colombia, Ecuador, and Bolivia (^{Smith, 2016}).

Locality records: Ahuachapán: Parque Nacional El Imposible (Puerta Mayor [Sector San Francisco Menéndez]) (^{Cave, 1983: 152}) (Fig. 29).

Years of collection: 1979.

Months of collection: June.

Topographic zone: coastal plain.

Elevation range: 250 m.

Comments: adults were found on or flying around the flowers of C. aculeata (^{Cave, 1983}).

Trigonopeltastes sallaei sallaei Bates, 1889

Distribution: Mexico, Guatemala, El Salvador, Honduras, Nicaragua, and Costa Rica (^{Smith, 2016}). Locality records: Ahuachapán: Parque Nacional El Imposible (Cerro de Loma de Paja [Sector San Francisco Menéndez]) (^{Cave, 1983: 152}); La Unión: Volcán de Conchagua (^{Howden, 1968: 38}) (Fig. 29).

Years of collection: 1979.

Months of collection: June.

Topographic zone: coastal mountains, coastal plain.

Elevation range: 250-1,190 m.

Comments: it is likely that the year and month of collection for the specimen from La Unión is the same (May 1958) as that of Trigonopeltastes variabilis Howden, since both species are reported in the same paper (^{Howden, 1968}). Adults were found on or flying around the flowers of C. aculeata (^{Cave, 1983}).

Trigonopeltastes variabilis Howden, 1968

Distribution: Mexico, Guatemala, El Salvador, and Honduras (^{Smith, 2016}).

Locality records: La Unión: Volcán de Conchagua (^{Howden, 1968: 49}) (Fig. 29).

Years of collection: 1958.

Months of collection: May.

Topographic zone: coastal mountains.

Elevation range: 1,190 m.